Three years ago I had a short and painful trip to China to meet my new colleague and friend Yongli Zhang (Department of Geology, Northeastern University, Shenyang). The China part was great; the pain was from an unfortunately-timed kidney stone I brought with me. Nevertheless, I got to meet my new colleagues and we continued on a project involving hard substrates in the Upper Carboniferous of north China. Above is one of our most important fossils, a chaetetid demosponge from the Benxi Formation (Moscovian) exposed in the Benxi area of eastern Liaoning Province. We are looking at a polished cross-section through a limestone showing the tubular, encrusting chaetetids. This month the paper on these fossils has at last appeared.
This closer view shows two chaetetids. The bottom specimen grew first, was covered by calcareous sediment, and then the system was cemented on the seafloor. After a bit of erosion (marked by the gray surface cutting across the image two-thirds of the way up), another chaetetid grew across what was then a hardground that partially truncated the first chaetetid. This little scenario was repeated numerous times in this limestone, producing a kind of bindstone with the chaetetids as a common framework builder.
Here is the closest view of the chaetetids, showing the tubules running vertically, each with a series of small diaphragms as horizontal floors.
Last week’s fossil was a chaetetid, introducing the group. They are hyper-calcified demosponges, and the classification of the fossil forms is still not clear. Their value for paleoecological studies, though, is clear. This particular chaetetid from the Benxi Formation preferred a shallow, warm, carbonate environment, and it was part of a diverse community of corals, fusulinids, foraminiferans, brachiopods, crinoids, bryozoans, gastropods, and algae. Such hard substrate communities are not well known in the Carboniferous, and this is one of the best.
Gong, E.P, Zhang, Y.L., Guan, C.Q. and Chen, X.H. 2012. The Carboniferous reefs in China. Journal of Palaeogeography 1: 27-42.
West, R.R. 2011a. Part E, Revised, Volume 4, Chapter 2A: Introduction to the fossil hypercalcified chaetetid-type Porifera (Demospongiae). Treatise Online 20: 1–79.
West, R.R. 2011b. Part E, Revised, Volume 4, Chapter 2C: Classification of the fossil and living hypercalcified chaetetid-type Porifera (Demospongiae). Treatise Online 22: 1–24.
Zhang, Y.L., Gong, E.P., Wilson, M.A., Guan, C.Q., Sun, B.L. and Chang, H.L. 2009. Paleoecology of a Pennsylvanian encrusting colonial rugose coral in South Guizhou, China. Palaeogeography, Palaeoclimatology, Palaeoecology 280: 507-516.
Zhang, Y.L., Gong, E.P., Wilson, M.A., Guan, C.Q.. and Sun, B.L. 2010. A large coral reef in the Pennsylvanian of Ziyun County, Guizhou (South China): The substrate and initial colonization environment of reef-building corals. Journal of Asian Earth Sciences 37: 335-349.
Zhang, Y., Gong, E., Wilson, M.A., Guan, C., Chen, X., Huang, W., Wang, D. and Miao, Z. 2017. Palaeoecology of Late Carboniferous encrusting chaetetids in North China. Palaeobiodiversity and Palaeoenvironments https://doi.org/10.1007/s12549-017-0300-5