Wooster’s Fossil of the Week: A scaphitid ammonite (Late Cretaceous of Mississippi)

February 24th, 2017

The beauty above is Discoscaphites iris (Conrad, 1858) from the Owl Creek Formation of Ripley, Mississippi. Megan Innis and I collected it during our expedition to the Cretaceous-Paleogene boundary in the southern United States last summer. It is a significant index fossil in biostratigraphy: the Discoscaphites iris Zone is the latest in the Cretaceous (the late Maastrichtian Stage). This animal lived in the final days of the Mesozoic Era just before the mass extinction 65.5 million years ago.

Discoscaphites iris is an ammonite, a type of extinct cephalopod mollusk related to the modern octopus, squid and nautilus. It had a planispirally-coiled shell with chambers divided from each other by complexly-folded walls. If you look closely near the top of the fossil above, you will see where the shell has flaked away revealing an internal mold of sediment and a peek at the folded walls inside. “Ammonite”, by the way, is a very old term for these fossils. Pliny the Elder himself used a variant of the name, which comes from the Egyptian god Amun with his occasional coiled ram’s horn headgear.

Reconstruction of an ammonite by Arthur Weasley (via Wikipedia).

Ammonite shells were made of the carbonate mineral aragonite. This is the mineral that makes many modern mollusk shells have prismatic colors, which we call nacreous. You may know it best as “mother of pearl” or as pearls themselves. Aragonite has an unstable crystal structure and so is not common in rocks older than a few million years. The original aragonite in our ammonite fossil is thus a bonus.

In an oddly topical note, Discoscaphites iris was recently found in the Upper Cretaceous of Libya, giving it a disjunct range from the US Gulf and Atlantic coasts to the Mediterranean coast of northern Africa (Machalski et al., 2009).


Machalski, M., Jagt, J.W.M., Landman, N.H. and Uberna, J., 2009. First record of the North American scaphitid ammonite Discoscaphites iris from the upper Maastrichtian of Libya. N. Jb. Geol. Paläont. Abh. 254: 373-378.

[Originally published April 24, 2011]

Wooster’s Fossil of the Week: A stromatoporoid (Middle Devonian of central Ohio)

February 17th, 2017

Stromatoporoids are very common fossils in the Silurian and Devonian of Ohio and Indiana, especially in carbonate rocks like the Columbus Limestone (from which the above specimen was collected). Wooster geologists encountered them frequently on our Estonia expeditions in the last few years, and we worked with at least their functional equivalents in the Jurassic of Israel (Wilson et al., 2008).

For their abundance, though, stromatoporoids still are a bit mysterious. We know for sure that they were marine animals of some kind, and they formed reefs in clear, warm seas rich in calcium carbonate (DaSilva et al., 2011). Because of this tropical habit, early workers believed they were some kind of coral, but now most paleontologists believe they were sponges. Stromatoporoids appear in the Ordovician and are abundant into the Early Carboniferous. The group seems to disappear until the Mesozoic, when they again become common with the same form and life habits lasting until extinction in the Late Cretaceous (Stearn et al., 1999).

The typical stromatoporoid has a thick skeleton of calcite with horizontal laminae, vertical pillars, mounds on the upper surface called mamelons, and dendritic canals called astrorhizae shallowly inscribed on the mamelons. These astrorhizae are the key to deciphering what the stromatoproids. They are very similar to those on modern hard sponges called sclerosponges. Stromatoporoids appear to be a kind of sclerosponge with a few significant differences (like a calcitic instead of an aragonitic skeleton).

Stromatoporoid anatomy from Boardman et al. (1987).

Top surface of a stromatoporoid from the Columbus Limestone showing the mamelons.

There is considerable debate about whether the Paleozoic stromatoporoids are really ancestral to the Mesozoic versions. There may instead be some kind of evolutionary convergence between two groups of hard sponges. The arguments are usually at the microscopic level!

The stromatoporoids were originally named by Nicholson and Murie in 1878. This gives us a chance to introduce another 19th Century paleontologist whose name we often see on common fossil taxa: Henry Alleyne Nicholson (1844-1899). Nicholson was a biologist and geologist born in England and educated in Germany and Scotland. He was an accomplished writer, authoring several popular textbooks, and a spectacular artist of the natural world. Nicholson taught in many universities in Canada and Great Britain, finally ending his career as Regius Professor of Natural History at the University of Aberdeen.

Henry Alleyne Nicholson (1844-1899) from the University of Aberdeen museum website.


Boardman, R.S., Cheetham, A.H. and Rowell, A.J. 1987. Fossil Invertebrates. Wiley Publishers. 728 pages.

DaSilva, A., Kershaw, S. and Boulvain, F. 2011. Stromatoporoid palaeoecology in the Frasnian (Upper Devonian) Belgian platform, and its applications in interpretation of carbonate platform environments. Palaeontology 54: 883–905.

Nicholson, H.A. and Murie, J. 1878. On the minute structure of Stromatopora and its allies. Linnean Society, Journal of Zoology 14: 187-246.

Stearn, C.W., Webby, B.D., Nestor, H. and Stock, C.W. 1999. Revised classification and terminology of Palaeozoic stromatoporoids. Acta Palaeontologica Polonica 44: 1-70.

Wilson, M.A., Feldman, H.R., Bowen, J.C. and Avni, Y. 2008. A new equatorial, very shallow marine sclerozoan fauna from the Middle Jurassic (late Callovian) of southern Israel. Palaeogeography, Palaeoclimatology, Palaeoecology 263: 24-29.

[Originally published on October 30, 2011]

Wooster’s Fossil of the Week: A receptaculitid (Middle Ordovician of Missouri)

February 10th, 2017

This week’s fossil is a long-standing paleontological mystery. Above is a receptaculitid from the Kimmswick Limestone (Middle Ordovician) near Ozora, Missouri. I think I found it on a field trip with Frank Koucky in the distant mists of my student days at Wooster, but so many outcrops, so many fossils …

Below is a nineteenth century illustration of a typical receptaculitid fossil. They are sometimes called “sunflower corals” because they look a bit like the swirl of seeds in the center of a sunflower. They were certainly not corals, though, or probably any other kind of animal. Receptaculitids appeared in the Ordovician and went extinct in the Permian, so they were confined to the Paleozoic Era. Receptaculitids were bag-like in form with the outside made of mineralized pillars (meroms) with square or diamond-shaped heads. The fossils are usually flattened disks because they were compressed by burial. You may notice now that the fossil at the top of this post is a mold of the original with the dissolved pillars represented by open holes. (Paleontologists can argue if this is an external or internal mold.)So what were the receptaculitids? When I was a student we called them a kind of sponge, something like a successor of the Cambrian archaeocyathids. In the 1980s a convincing case was made that they were instead a kind of alga of the Dasycladales. Now the most popular answer is that they belong to that fascinating group “Problematica”, meaning we have no idea what they were! (Nitecki et al., 1999). It’s those odd meroms that are the problem — they appear in no other known group, fossil or recent.

I find it deeply comforting that we still have plenty of fossils in the Problematica. We will always have mysteries to puzzle over.
Another Wooster receptaculitid specimen, this time seen from the underside showing side-views of the meroms.
Diagram of a receptaculitid in roughly life position showing its inflated nature and pillar-like meroms. From Dawson (1880, fig. 25): a, Aperture (probably imaginary here). b, Inner wall. c, Outer wall. n, Nucleus, or primary chamber. v, Internal cavity.

Finally, this is what a typical receptaculitid looks like in the field (Ordovician of Estonia). Note that nice sunflower spiral of the merom ends.


Dawson, J.W. 1880. The chain of life in geological time: A sketch of the origin and succession of animals and plants. The Religious Tract Society, 272 pages.

Nitecki, M.H., Mutvei, H. and Nitecki, D.V. 1999. Receptaculitids: A Phylogenetic Debate on a Problematic Fossil Taxon. Kluwer Academic/Plenum, 241 pages.

[Originally published on September 18, 2011]

Wooster’s Fossils of the Week: Peanut worms from the Silurian of Illinois

February 3rd, 2017

1-lecthaylus-gregarius-5-copyThis week’s fossils are a set of cool sipunculan worms from the Lockport Shale Member of the Racine Formation (Wenlockian, Silurian) of Blue Island, Illinois (which, it turns out, is not an island.). This is Lecthaylus gregarius Weller, 1925. (There is a common misspelling of the genus name as “Lecathylus”, which is how it is labeled in our collection.) They are masses of partially-carbonized bodies and external molds in a very fine-grained matrix. They are well known from this particular fossil-lagerstätte (a fossil fauna of remarkable preservation) in northern Illinois.

The Phylum Sipuncula did not often make it into the fossil record because of their entirely soft bodies, but a few are preserved way back in the Cambrian Chengjiang and Burgess Shale faunas. They show virtually no evolutionary changes in their long run to today, at least not in their outer form. They are commonly known as “peanut worms”.

2-lecthaylus-gregarius-2This is an example of the preservation modes: a black carbon film that has mostly flaked away, leaving behind a detailed external mold of the squashed peanut worms.

3-lecthaylus-gregarius-1Sipunculan bodies are divided into a main thick posterior trunk and a narrow, retractable anterior “introvert”. We’re looking here at the anterior introvert of Lecthaylus gregarius.

4-lecthaylus-gregarius-3-copyThis is the squat trunk of Lecthaylus gregarius.

5-themiste_petricola_evertedHere is the modern sipunculan Themiste petricola with introvert extended. It is the same basic plan as the Silurian Lecthaylus gregarius. Image from Wikipedia courtesy of Tomás Lombardo and Guillermo A. Blanco.

6-themiste_petricola_invertedThe modern sipunculan Themiste petricola with its introvert retracted. Image from Wikipedia courtesy of Tomás Lombardo and Guillermo A. Blanco.

stuart-weller-1870-1927Lecthaylus gregarius was described and named by Stuart Weller (1870-1927), an American paleontologist and geologist. He was born in the small town of Maine, New York. He earned a Bachelor’s degree in geology at Cornell University in 1894 followed by a PhD at Yale in 1901. Shortly after his Cornell degree, though, Weller traveled to the University of Chicago, where he worked his way through the ranks from a research associate to a full professor of Paleontology and Geology in 1915. He was also the director of the Walker Museum at the University of Chicago, and in 1926 he was president of the Paleontological Society. One of his sons, J. Marvin Weller (1899-1976) had a remarkably similar career as a stratigrapher and paleontologist.


Kluessendorf, J. 1994. Predictability of Silurian Fossil‐Konservat‐Lagerstatten in North America. Lethaia 27: 337-344.

Roy, S.K. and Croneis, C. 1931. A Silurian worm and associated fauna. Field Museum of Natural History, Geological Series IV(7): 229-247.

Weller, S. 1925. A new type of Silurian worm. Journal of Geology 33: 540-544.

Wooster’s Fossil of the Week: Ammonite septa from the Upper Cretaceous of South Dakota

January 27th, 2017

This week we have an ammonite from the Pierre Shale (Upper Cretaceous, Campanian-Maastrichtian) of southwestern South Dakota. It was collected on a wonderful field expedition in June 2008 with my friend Paul Taylor (The Natural History Museum, London) and my student John Sime. Ammonites are extremely common in this interval, but I like this one because it is broken in such a way to expose its complex internal walls, called septa. We are looking at a cross-section of a coiled ammonite showing an early whorl in the upper left surrounded by a later whorl. The septa are fluted at their margins as they meet the outer wall. The wiggly boundary line between a septum and the outer wall is called a suture.

Ammonite septa are remarkably complex, showing fractal patterns. Why did these animals, extinct for 66 million years, evolve such complexity in their septa? This is a hotly debated topic in paleontology. The most popular explanations include strengthening the walls of the shell to resist hydrostatic pressure at depth, buttressing the shell against the crushing pressures of biting predators, and increasing soft-tissue (mantle) surface areas for physiological advantages. Klug and Hoffman (2015) have an excellent summary of these ideas. Lemanis et al. (2016) have a fascinating mathematical study that suggests the answer in many cases complex sutures “seem to increase resistance to point loads, such as would be from predators.”

The astonishing English polymath Robert Hooke (1635-1703) took considerable interest in ammonites and their complicated septa. We have no contemporary images of him, but based on descriptions, Rita Greer painted the above portrait in 2004. Hooke’s life was as complex as the suture patterns he studied, so I leave you to other sources on him. Note in the portrait above, though, the ammonite!

These are drawings by Robert Hooke of ammonites and their suture patterns (from Kusukawa, 2013). It is a single image mirror-reversed. Beautiful.


Derham W. 1726. Philosophical experiments and observations of the late eminent Dr. Robert Hooke, S.R.S. and Geom. Prof. Gresh., and other eminent virtuoso’s in his time. London: Derham.

Garcia-Ruiz, J.M., Checa, A. and Rivas, P. 1990. On the origin of ammonite sutures. Paleobiology 16: 349-354.

Klug, C. and Hoffmann, R. 2015. Ammonoid septa and sutures. In: Ammonoid Paleobiology: From anatomy to ecology (p. 45-90). Springer Netherlands.

Kusukawa, S. 2013. Drawings of fossils by Robert Hooke and Richard Waller. Notes Rec. R. Soc., 67: 123-138.

Lemanis, R., Zachow, S. and Hoffmann, R. 2016. Comparative cephalopod shell strength and the role of septum morphology on stress distribution. PeerJ 4:e2434

Wooster’s Fossils of the Week: Revisiting a pair of hyoliths from the Middle Ordovician of Estonia

January 20th, 2017

We met these modest internal molds of the mysterious hyoliths about five years ago. With a dramatic new development in hyolith studies, they are worth seeing again.

These fossils are internal molds (the sediment that filled the shell) of of flattened cones composed of the carbonate mineral aragonite. The aragonite shells dissolved away after burial, leaving the cemented sediment behind. That’s what we see above, in their stark simplicity. (We also see wiggly indentations that are the trace fossil Arachnostega, which is what I collected them for in the first place.) They were found in the Middle Ordovician of Estonia.

Hyalites, though common throughout the Paleozoic, have been difficult to place in a taxonomic category. Because of their easily-dissolved aragonite skeletons, most fossils are like these — simple molds and casts. A few were found with some preserved internal organs, which added to the intrigue. Their flattened conical shells had a hinged lid (operculum) over the open end. Extending from each side in the space between the operculum and cone were two calcareous rods called helens (a name deliberately chosen so as not to evoke a particular function). They were rumored to be deposit-feeders, based on no real evidence, it turns out.

An excellent paper appeared earlier this month showing dramatic evidence of hyolith soft parts in the Cambrian of western Canada (Moysiuk et al., 2017). The authors reconstruct the iconic Cambrian hyolith Haplophrentis “as a semi-sessile, epibenthic suspension feeder that could use its helens to elevate its tubular body above the sea floor”. Their primary evidence is a magnificently preserved lophophore (tentacular filter-feeding apparatus) and a U-shaped digestive tract with a dorsolateral anus. These features not only give the hyoliths a life mode and feeding habit, they place them systematically among the lophophorates, a group that includes brachiopods, phoronids and bryozoans.

Haplophrentis in the Burgess Shale (Middle Cambrian) at the Walcott Quarry, Burgess Pass, British Columbia, Canada.

Reconstruction of Haplophrentis on the Cambrian sea floor. The tentacular lophophore is seen extending out underneath the operculum. Beautiful art by D. Dufault of the Royal Ontario Museum.

It’s not often we see such dramatic changes in the taxonomic placement and paleoecological habits of a large, extinct group. It is also not often that invertebrate fossils make headlines!


Moysiuk, J., Smith, M.R. and Caron, J.B. 2017. Hyoliths are Palaeozoic lophophorates. Nature doi:10.1038/nature20804

Wooster’s Fossils of the Week: New review paper on architectural design of trace fossils

January 13th, 2017

screen-shot-2016-12-04-at-2-59-29-pmLast year my friend Luis Buatois led a massive project to review essentially all trace fossil invertebrate ichnogenera (523!) to place them in a series architectural design categories (79). This is a new way to assess patterns of ichnodisparity (variability in morphology of trace fossils). I was proud to have a role in this work, along with Max Wisshak and Gabriela Mángano. The paper has now appeared in Earth-Science Reviews (Buatois et al., 2017).

My contributions were mostly with the bioerosion traces (along with Max), so I show Figure 65 from the paper above. Its caption: Examples of pouch borings (Category 65). A: Petroxestes pera, Ordovician, Whitewater Formation, Ohio, USA. B: Rogerella isp. in a belemnite rostrum. Jurassic, Spain. C: SEM of Rogerella isp. in an epoxy resin cast of an Echinocorys echinoid test. Upper Cretaceous, Palm Bay, Thanet, Kent, UK. D: Umbichnus inopinatus in a bivalve shell. Lower Pliocene, Huelva, Spain. Photograph courtesy of Jordi Martinell. E: SEM of Aurimorpha varia in an epoxy resin cast, including the holotype in the upper right. Middle Pennsylvanian, Desmoinesian, Boggy Formation, Buckhorn Asphalt Quarry, Oklahoma, USA.

The abstract of the paper explains the work and our ambitions for it: Ichnodisparity has been recently introduced as a concept to assess the variability of morphologic plans in biogenic structures, revealing major innovations in body plan, locomotory system and/or behavioral program. Whereas ichnodiversity is measured in terms of the number of ichnotaxa (i.e. ichnogenera or ichnospecies), ichnodisparity is evaluated based on the identification of categories of architectural design. Seventy-nine categories of architectural designs (58 for bioturbation structures and 21 for bioerosion structures), encompassing 523 ichnogenera (417 for bioturbation structures and 106 for bioerosion structures), are defined. They are restricted to invertebrate ichnotaxa, whereas vertebrate trace fossils were not included. Although the scheme is designed to be comprehensive, the proposed categories are necessarily works in progress because of the state of flux in ichnotaxonomy and the need to adjust the definitions of categories according to the scope and scale of the analysis. Although it may be said that the establishment of categories of architectural design is to a certain degree a subjective enterprise, this is not different from ichnotaxonomy because classifying trace fossils from a taxonomic perspective implies observing the morphology of the trace and interpreting it in terms of behavior. The concept of ichnodisparity is free of some of the vagaries involved in ichnotaxonomy. The fact that ichnodiversity and ichnodisparity exhibit different trajectories during the Phanerozoic underscores the importance of adding the latter to the ichnologic toolkit.
screen-shot-2016-12-04-at-3-03-44-pmFigure 80 above contrasts ichnodisparity and ichnodiversity. The five different ichnogenera illustrated in the upper portion of the diagram represent minor variations of the same architectural design. The lower portion of the diagram represents the same ichnodiversity level, but with a much higher ichnodisparity. The two hypothetical situations bear different implications regarding the extent of evolutionary innovations.

We hope that this work is long useful in paleontology, especially for projects sorting out the evolution of invertebrate communities.


Buatois, L., Wisshak, M., Wilson, M.A. and Mángano, G. 2017. Categories of architectural designs in trace fossils: A measure of ichnodisparity. Earth-Science Reviews 164: 102-181.

Wooster’s Fossils of the Week: Upper Ordovician brachiopods and bryozoans from paleontology class collections

January 6th, 2017

1-geopetal-tommyLast semester the Invertebrate Paleontology class at Wooster had its annual field trip into the Upper Ordovician of southern Ohio. We had a great, if a bit muddy, time collecting fossils for each student’s semester-long project preparing, identifying, and interpreting their specimens. Like all research, especially when it starts in the field, there were discoveries and surprises. I always highlight a particular specimen collecting by a student in this blog.

Above is a cross-section of a specimen found by Tommy Peterson (’19). It is the rhynchonellid brachiopod Hiscobeccus capax almost completely enveloped by an encrusting trepostome bryozoan. We’ve cut through the center of the brachiopod, revealing gray micritic sediment and clear calcite crystals. We can infer from this simple specimen that the brachiopod died and its shell remained articulated. Sediment drifted in, filling the bottom half of the shell. The bryozoan eventually sealed it all up as it used the brachiopod shell for a hard substrate on a muddy seafloor. The remaining void space was filled in by the precipitation of calcite crystals. You can see that the crystals nucleated from the outer margin of the cavity and grew inwards, a kind of calcareous geode. I’m intrigued by the irregular sediment surface and the manner in which calcite nucleated upwards from it. I suspect this sediment was itself cemented before the calcite crystals appeared.

This kind of structure is called a geopetal. It shows the “way up” at the time of crystal formation. Gravity held the pocket of sediment in the bottom of the shell, leaving the void at the top. Nice little specimen.

2-constellaria-alexisThis star-studded bryozoan found by Alexis Lanier (’20) was going to be the Fossil of the Week, but then I saw that last year I highlighted the very same species! I think the bryozoan Constellaria is cool. Read all about it and its history at the link.

3-table-of-traysHere are the completed specimen trays for half the class. (Grading this project took, as you might imagine, considerable time!). Every week in lab, after we had done the assigned work, we got out the trays and cleaned, prepared, and identified the specimens. Students learned how to use the rock saws and make acetate peels of the bryozoans and corals.

4-tray-insideInside a typical tray. We are very grateful for the many online sources to aid identification of these Cincinnatian fossils. Three in particular were most valuable: Alycia Stigall’s Digital Atlas of Ordovician Life, Steve Holland’s stratigraphic and paleontological guide to the Cincinnatian, and the spectacular Dry Dredgers website.

Ohio is a paleontological paradise!

Wooster’s Fossils of the Week: Geological Magic Lantern Slides from the 19th Century (Part III)

December 16th, 2016

18-devonion-period[Note: Wooster’s Fossil of the Week is on holiday until January 2017.]

This is the last post illustrating the 19th Century Magic Lantern Slides recently found in Scovel Hall of Wooster’s Geology Department. Please see the December 2 post and the week before for details. To review, these slides are 4×8 inches with the image fixed on glass bolted into a thin slab of wood with metal rings. They are chromolithograph slides, each stamped “T.H. McAllister, Optician, N.Y.”. McAllister was the most prominent of many American producers of lantern slides in the late 19th century.

This last set of slides in our collection was apparently used in our old “Historical Geology” courses to evoke the geological time periods. The top image is simply labeled “Devonian“. The trees on the right appear to be towering lycopods, a kind of seedless vascular plant. They were common in the Devonian and are still around today. I can’t tell what the other plants are in the image. The rapid rise of large plants in the Middle Devonian has been called the “Devonian Explosion”. These early forests had significant effects on atmospheric composition, soil formation, erosion, and sediment transport.

[UPDATE: Please see the excellent comments by Ben Creisler. He has given us much new information and numerous links explaining the history of these images. I’ve left my amateur text in place only to record the original post! MW]

19-carboniferous-periodCarboniferous” is the title of this slide. It is dramatic, seemingly showing a Carboniferous forest dominated by ferns being torn apart by a swelling tide. Could this be a comment on the interbedding of marine and terrestrial rock units so common in the Upper Carboniferous of North America?

20-permian-periodFerns are again in the foreground of this Permian scene. I have no explanation for the mountainous seashore landscape, except that the red color of the rocks may represent the New Red Sandstone of Great Britain.

21-transition-periodThis slide is enigmatically labeled “Transition Period”. I suspect it represents the Triassic, a period just after the Permian and thus part of the transition into the Mesozoic. The shrubby plants in the foreground appear to be cycads with massive yellow cones emerging from their tops.

22-eeocen-periodThis image of the “Eocene” is the first of these period slides to depict animals (the herd of ungulates across the river and the bird in the foreground). This may mean these slides were meant to show the progression of plant life over geological time. The forests here look dominated by conifers and angiosperms.

23-miocene-periodThis is a “Miocene” image. I don’t know how I’d distinguish it from the Eocene view above.

24-drift-periodOur final slide shows what the “Drift Period”, which is clearly the Pleistocene. Not only do we have cave bears in the foreground and a herd of bison in the river, there seems to be a massive pile of ice in the left rear!

I have not discovered the artist responsible for these illustrations. If you know, please tell me in the comments!

[UPDATE: Please see excellent information and links by Ben Creisler in the comments below. Thanks, Ben!]


Wooster’s Fossils of the Week: Ordovician bioerosion trace fossils

December 9th, 2016

screen-shot-2016-12-03-at-2-06-03-pmThis week’s post is a celebration of the appearance of a remarkable two-volume work on trace fossils and evolution. The editors and major authors are my friends Gabriela Mángano and Luis Buatois (University of Saskatchewan). They are extraordinary geologists, paleontologists and ichnologists (specialists on trace fossils). They led this massive effort of multiple authors and thousands of manuscript pages. Turns out they are inspiring scientific leaders as well as sharp-eyed editors.

My contribution is in the first volume within a chapter (co-authored with Gabriela, Luis, and Mary Droser of the University of California, Riverside) entitled “The Great Ordovician Biodiversification event”. We examine here the relationship between trace fossils and the critical evolution of marine communities through the Ordovician. My main responsibility was sorting out the changes in the bioeroders over the course of the period. Way back in 2001, Tim Palmer and I noticed a rise in bioerosion trace fossil diversity and abundance in the Middle and Late Ordovician. We grandly called it the “Ordovician Bioerosion Revolution”. The concept and name stuck.

The top image is Fig. 4.8 from the book. The caption: Upper Ordovician bioerosion structures. (a) Trypanites weisi (cross-sectional view) in a carbonate hardground. Katian, Grant Lake Limestone, near Washington, Kentucky, USA; (b) Trypanites weisi (bedding-plane view) in a carbonate hardground. Katian, Grant Lake Limestone, near Manchester, Ohio, USA; (c) Palaeosabella isp. in a trepostome bryozoan. Katian, Whitewater Formation, near Richmond, Indiana, USA; (d) Petroxestes pera. Katian, Whitewater Formation, Caesar Creek Lake emergency spillway, near Waynesville, Ohio, USA; (e) Ropalonaria venosa in a strophomenid brachiopod. Katian, Liberty Formation near Brookville, Indiana, USA.

screen-shot-2016-12-03-at-2-08-42-pmThe cover of the book, which is described here on the publisher’s website.


Mángano, G., Buatois, L., Wilson, M.A. and Droser, M. 2016. The Great Ordovician Biodiversification event, p. 127-156. In: Mángano, G. and Buatois, L. (eds.), The trace-fossil record of major evolutionary events. Topics in Geobiology 39 (Springer).

Wilson, M..A. and Palmer, T.J. 2001. The Ordovician Bioerosion Revolution. Geological Society of America Annual Meeting, Boston, Paper No. 104-0. November 7, 2001.

Wilson, M.A. and Palmer, T.J. 2006. Patterns and processes in the Ordovician Bioerosion Revolution. Ichnos 13: 109-112.

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