Wooster’s Fossil of the Week: A grazed oyster from the Middle Jurassic of Gloucestershire, England

March 24th, 2013

Praeexogyra_acuminata_585This small oyster is not in itself unusual. In fact, it is one of the most common fossils in the Jurassic of western Europe: Praeexogyra acuminata (Sowerby, 1816). It may be better known by its older name: Ostrea acuminata. Local collectors call it the “sickle oyster” because of its curved shape. This specimen is from the Sharp’s Hill Formation (Middle Bathonian) exposed in the Snowshill Quarry near Moreton-in-Marsh, Gloucestershire, England. I collected it on my first trip to England in 1985.
Praeexogyra_acuminata_closerWhat attracted me to this particular shell can be seen in the above close-up: lots of little straight lines incised across its outer surface (along with a serpulid worm tube). The lines were scraped by the Aristotle’s Lantern of one or more regular echinoids (sea urchins). This is the trace fossil Gnathichnus pentax Bromley, 1975. We met this fossil last month cut into a Cretaceous oyster from Israel. One or more echinoids grazed over this Jurassic oyster, probably consuming algae and other organic materials.

Praeexogyra acuminata was an epifaunal filter-feeder, meaning it lived on the substrate sucking in seawater and sorting from it organic material for food. During the Middle Jurassic these oysters were so common that their shells formed thick deposits. It is possible that some deposits rich in these shells were formed in brackish waters rather than under fully marine conditions.

Ostrea acuminata was named by by the enthusiastic English natural historian James Sowerby (1757-1822). We met him earlier as the author of a Cretaceous bivalve genus.

References:

Bernard-Dumanois, A. and Delance, J-H. 1983. Microperforations par algues et champignons sur les coquilles des «Marnes à Ostrea acuminata (Bajocien supérieur) de Bourgogne (France), relations avec le milieu et utilisation paléobathymétrique. Geobios 16: 419-429.

Bernard-Dumanois, A. and Rat, P. 1983. Etagement des milieux sédimentaires marins. Paléoécologie des Huîtres dans les “Marnes à Ostrea acuminata” du Bajocien de Bourgogne (France). Comptes rendus de l’Académie des sciences Paris 296: 733-737.

Hudson, J.D. and Palmer, T.J. 1976. A euryhaline oyster from the Middle Jurassic and the origin of true oysters. Palaeontology 19: 79-93.

Wooster Geologist at Valley Forge, Pennsylvania

March 20th, 2013

ValleyForgeHuts032013BRYN MAWR, PENNSYLVANIA–While visiting my friends and colleagues Katherine and Pedro Marenco at Bryn Mawr College, I visited the nearby Valley Forge National Historical Park. Everyone will remember, of course that this is the place outside Philadelphia that the Continental Army made its rough winter quarters in 1777-1778. The huts above are reconstructions of the soldiers’ quarters on the windy and cold fields. Commander-in-Chief George Washington chose this place because it was easily defensible, had plenty of timber for construction and fuel, and was close enough to British-occupied Philadelphia to keep an eye on the enemy — yet not so close to be likely attacked.

LedgerOutcrop032013

As a geologist, of course, I also looked for the rocky bones beneath the landscape. They were easily found in the above cliff near the main parking area. This is the Ledger Dolomite, a Cambrian unit found throughout this part of eastern Pennsylvania.

LedgerDolomite032013The Ledger Dolomite here is distinguished by these fine laminations visible on its weathered cross-sections. These are apparently stromatolites: laminar structures built by bacterial mats. We’ve met Cambrian stromatolites before in this blog.

Smilodon_gracilisI was surprised to learn that there is also a significant middle Pleistocene fossil deposit in Valley Forge called the Port Kennedy Bone Cave. This is a sinkhole deposit within the Ledger Dolomite. A particularly large sinkhole apparently trapped a variety of animals, including the gracile sabre-tooth Smilodon gracilis, the skull of which is on display in the Valley Forge Historical Park visitor center. S. gracilis was the smallest and earliest member of its genus. The Port Kennedy Bone Cave was one of the first fossil assemblages that the famous paleontologist Edward Drinker Cope studied. The location was lost to science until its rediscovery in 2005.

ValleyForgeCannon032013This is the requisite cannon image, even though no battle was fought here. It is nevertheless a dramatic place for the privations the soldiers suffered during the darkest days of the Revolutionary War. It is hard to imagine the conditions in 1777-1778 now since highways and casinos surround the old encampment.

Wooster’s Fossil of the Week: Cast of a lower jawbone of the largest ape ever (Pleistocene, southern China)

March 17th, 2013

Gigantopithecus_blacki_mandible_010112The above is one of my favorite “fossils”, a commercially-available cast of the lower jawbone of Gigantopithecus blacki, a giant extinct ape. It was produced from an actual Pleistocene fossil found in a cave near Liucheng, Guangxi, in southern China. I like it especially because it is sometimes associated with the mythical “Bigfoot”.

Gigantopithecus blacki was the largest ape that ever lived: up to three meters tall and weighing over 500 kilograms. (G. blacki is known only from teeth and mandibles such as that shown above, so these size estimates are based on scaling.) It was a contemporary with early versions of our own species, which must have led to a few astounding encounters for our ancestors. G. blacki was two or three times heavier than the largest gorillas today.

Gigantopithecus blacki appears to have lived in bamboo forests. Striations on its teeth, and the occasional phytolith stuck in the enamel, shows that this species was a vegetarian. It may have even had a lifestyle much like today’s pandas.

The molars of Gigantopithecus blacki look surprisingly like ours with their multiple cusps and broad surfaces. This is the result of convergent evolution and not an indication of a recent common ancestry. (They are analogous features, not homologous.) G. blacki is now classified in the Subfamily Ponginae with their cousins the orangutans.

What is most fun about Gigantopithecus these days is its association with the “Bigfoot” illusion. Look at how seriously the people at the “Bigfoot Field Researchers Organization” take the possible connection of Gigantopithecus and Bigfoot. Despite their objections, we really can wonder why we’ve never found evidence of this giant ape in North America, including bones, teeth, legitimate footprints or real photographs. A living three-meter tall ape is a bit difficult for science to have missed! (Unless, of course, Bigfoot has supernatural powers.)

References:

Coichon, R. 1991. The ape that was – Asian fossils reveal humanity’s giant cousin. Natural History 100: 54–62.

Ciochon, R., et al. 1996. Dated co-occurrence of Homo erectus and Gigantopithecus from Tham Khuyen Cave, Vietnam. Proceedings of the National Academy of Sciences of the United States of America 93: 3016–3020.

Jin, C., et al. 2009. A newly discovered Gigantopithecus fauna from Sanhe Cave, Chongzuo, Guangxi, South China. Chinese Science Bulletin 54: 788-797.

Last day in the Mojave Desert for Wooster Geologists … this year

March 15th, 2013

RedRocks031513LAS VEGAS, NEVADA–We left the fine people at the Desert Studies Center in Zzyzx in the morning and drove in an all-too-irregular way through Las Vegas (sorry about that, other drivers!) to the spectacular Red Rocks National Conservation Area northwest of the city. Here we see a Paleozoic section thrust from the west over the white and red rocks of the Aztec Sandstone (Lower Jurassic). The view above is looking directly west from the visitor center. I don’t know all the names of these peaks, but I do recognize Mount Wilson on the left. (I did my dissertation in the Carboniferous rocks exposed in the background, but I don’t think they named the mountain after me.)

Our students, faculty and staff scattered for a couple of hours in this park on yet again a very pleasant day. Many of them climbed the sandstone cliffs, no doubt impressed by the large foresets showing that these sediments were deposited in massive dunes blown in from the east.

RedRocksVisitorCenter031513The redesigned visitor center at Red Rocks is excellent. Above is a view of part of the outside portion that takes advantage of the sun’s angle and cooling breezes. It includes a desert tortoise enclosure where we saw three of these endangered animals. This complex is so attractive and efficient that Yoav Avni took many photographs so that he could show Israeli planners some new ideas for their public displays.

At noon we drove back into Las Vegas and the open jaws of the McCarran Airport Complex. We turned in all four vans with no problems, got our boarding passes for our various destinations, and now are simply waiting in the airport for our flights out. An excellent and enjoyable trip in all respects. Here’s to Jason and Rob at the Desert Studies Center for their friendly and efficient management, to the rangers of the National Park Service for their helpful advice, to the faculty and staff leaders and drivers (so many miles!), and finally to the students who could not have been better ambassadors for Wooster Geology. Their enthusiasm and good humor is what makes these trips so special. We will return!

Exploring the Tecopa region … and finally the trilobites!

March 14th, 2013

SaltSpring031413ZZYZX, CALIFORNIA–This was our last full day of geological adventures in the Mojave Desert for 2013. The weather was even warmer than yesterday, with the temperature in Baker at 4:00 p.m. a delightful 94°F. We began in the usual way from our temporary home at the Desert Studies Center, heading north of Baker again to Shoshone and the Tecopa area. Our first stop was at Salt Creek Spring in the Salt Spring Hills, one of the rare riparian environments in this part of the desert. This place is also notable as the first place gold was found in southern California. This discovery in 1849 began the boom in mining throughout the Mojave Desert.

TafoniMelissa031413One of the interesting geological features we noted was the occurrence of the familiar tafoni, but this time in a deeply eroded outcrop of quartz monzonite (shown above with Melissa Torma). Our companion Yoav Avni believes this may represent a Pleistocene pluvial phase of erosion that has been exhumed in the modern landscape. Tafoni features in granite would seem to require more water than is present today in this system, thus placing these features during a wet interval makes sense.

ShoshoneMuseum031413We next returned to the little settlement of Shoshone, this time visiting the town museum (above). It is well worth touring for its local artifacts, some wildlife displays, and the bones and teeth of the Shoshone Mammoth found in the Tecopa lake sediments.

Archidiskodon_tamenensis031413The mammoth (Archidiskodon tamenensis on the sign) has been very well preserved by the lake clays and silts, especially the diagnostic teeth and jaws. It is an impressive find and an important addition to our knowledge of the ancient Lake Tecopa ecosystem.

OliviaBrownResting Springs031413The Resting Springs Welded Tuff exposed east of Shoshone is one of the most famous roadcuts in the Mojave Desert. Above we see Olivia Brown exploring the glassy black middle of the unit. The colors and hardnesses of the rocks here show degrees of welding from the intense heat developed in this pyroclastic flow. It is technically a devitrified pumice tuff, welded tuff, and vesicular vitrophyre dated by K-Ar methods as 9.5 million years old. The fiamme are fantastic.

JonahDragFold031413In the same outcrop of the Resting Springs Tuff are some impressive faults. Jonah Novek is showing above how he knows this is a normal fault because the drag fold shows downward movement of the hanging wall (the side he is standing on).

TrilobiteCollecting031413At lunchtime we arrived at Emigrant Pass east of Tecopa to collect trilobites from the Cambrian Carrara Formation exposed on the north side of the highway (which happened to be the Old Spanish Trail). After a bit of exploring, our ace field geologist Shelley Judge found the most fossiliferous outcrops. We found hundreds of the fascinating critters, leaving most for future field trips. I like the view above, by the way, because it shows Telescope Peak on the far distant skyline.

TrilobiteWhole031413The best trilobite of the day (and there were many good specimens) was collected by Olivia Brown. It is a rare whole specimen, apparently of the genus Olenellus. You can see all the thoracic segments and spines, as well as the pygidium. The way the cephalon is separated from the thorax may indicate that it is a preserved exuvium (molt). Very nice.

ChinaRanchSign031811aFollowing a geological tradition transferred from Sonoma State University geologists to us via my friend Matthew James, we visited China Ranch south of Tecopa and had date milkshakes. Delicious and refreshingly cool on what turned out to be a very hot day. We also had a good geology lesson with the fanglomerates and ancient lakebeds exposed around the date groves.

TecopaPlaya031413While driving out of Tecopa, we stopped to walk out onto the Lake Tecopa playa, which was covered by a thin crust of salts. From the left are Meagen Pollock, Melissa Torma, Shelley Judge and Olivia Brown. We observed the modern playa sediments juxtaposed with the ancient lake strata exposed as low mounds.

WadeExit031413Last of all was a stop to briefly recognize the Harry Wade exit route out of Death Valley at an historical marker on the highway back to Baker. We came full circle at this point, since we started with Death Valley and now ended looking back into that source of such hardship in 1849.

Our final scientific event of the day was an excellent tour of the night sky by the Laboratory Technician of the Vassar College Environmental Sciences and Geography Department, Rick Jones. The sky is very dark and inviting in this lonely part of the desert.

Enjoying the geology of the Mojave National Preserve

March 13th, 2013

StudentsLavaCave031313ZZYZX, CALIFORNIA–Today the Wooster Geologists explored the Mojave National Preserve. It is a beautiful, spacious, diverse place well maintained and protected by the National Park Service. Our first stop of the morning is shown above. We explored a lava tube in the cinder cones portion of the preserve. We descended into the dark tunnel by the steps above, and then studied the walls and ceiling with our flashlights, as Melissa Torma is doing below.
MelissaCave031313Lava tubes are formed when a lava flow cools on the top and sides, and then the remaining lava flows out, leaving behind the empty shell. In this particular lava tube we could see various level markings the lava left on the walls as it drained away. We also saw spatter and “lavacicles” hanging from the ceiling much like icicles or stalactites in a cave. My favorite part is where we can see from below a granitic boulder that was caught up in the flowing lava.
CambrianLimestoneOutcrop031313South of the lava tube along Kelbaker Road is a small outcrop of Cambrian limestone. We examined these rocks for a short time (above) to sort out their lithologies and the paleoenvironment in which the sediment accumulated. The clues included extensive and diverse horizontal burrow systems (shown below) and numerous oncoids with shelly nuclei.
BurrowsLimestone031313The students determined that the burrows were filled with dolomite from later diagenesis of the sediment, and that the oncoids showed the system was deposited well within the photic zone. We think this rock is a biomicrite formed around the storm wavebase on a shallow carbonate platform.

KelsoDunes031313One of our favorite places is Kelso Dunes. These massive piles of sand are stabilized at their bases but still moving about at their crests. We’ve studied them so many times in the Desert Geology and Sedimentology & Stratigraphy courses that we needed to say little about them. The students raced to the top, noting the diverse sand composition (so much more than just quartz) and eolian structures on the way.

TafoniAnanda031313Ananda Menon is here showing us one of the magnificent tuff beds at the Hole in the Wall locality in the easternmost portion of the Mojave National Preserve. The holes in the wall (is that how the place was named?) are examples of tafoni, a rock weathering pattern. We hiked the Rings Loop Trail examining the tuff units and their diverse clasts, including pumice and charcoal. We also saw good examples of Indian petroglyphs of unknown age.

QtzMonzoniteCimaDome031313While crossing Cima Dome on the way back to Zzyzx, the group visited an outcrop of quartz monzonite which forms at least part of the intrusive body under the uplift. As with most of our outcrops, it was a story of both petrology and structural deformation.

AlienFreshJerky031313Finally, at the end of the day I could not resist taking the Wooster Geologists to the unique store in Baker called “Alien Fresh Jerky“. Half the group loved it, the other seemed offended by the epic cheesiness. At least Melissa Torma and Jonah Novek enjoyed meeting members of the Galactic Peace Patrol parked in front.

Folded rocks, volcanoes, and a little of the Old West

March 12th, 2013

CalicoMemberFolds031213ZZYZX, CALIFORNIA–The spectacular set of folds above are exposed in the lower parking lot of Calico Ghost Town near Barstow, California. This is a famous site that has been visited by hundreds of geologists, but there are still many mysteries about the causes of this deformation and even the correlation of these rocks in the region. The section makes up the Calico Member of the Barstow Formation, which is apparently below the exposed units of the Barstow Formation in Owl Canyon and Rainbow Basin we saw yesterday.
ShelleyCalico031213As a true structural geologist, Shelley Judge is here explaining the folding of these beds with her hands and, later, a sketch on notepaper. “Explaining” is not quite the right word: she wrangles information and observations out of the students first before developing the deformation hypotheses. It is not just about a bit of compression!
StudentsCalico031213After exploring a wadi west of the town, we then spent the rest of the morning in Calico. The students above (from the left, Kyle Burden, Steph Bosch, Sarah Frederick, Olivia Brown and Alex Hiatt) are enjoying the main street of this reconstructed 1880s mining community. There are lots of attractions here, from sarsaparilla in the saloon to a classic old-timey mystery shack. One gentleman, hearing that we were geologists, kindly explained to us how Calico Mountain is an extinct volcano with “sandy dikes” full of metals and remnants of lava flows on its sides. It was fun to see our students grapple with the question: do we correct this nonsense or just smile and avoid the hassle? (I just let it pass because I was hungry for lunch!)
AmboyCrater031213In the afternoon we drove south and east along Interstate 40 and Route 66 (yes, the Route 66) to Amboy Crater (shown above). The day had warmed considerably — my thermometer said it was now 94°F. Nevertheless, the stalwart geology crew walked to this gorgeous cinder cone and climbed to its rim from the inside (shown below). I enjoyed learning about the subtle features of the lava flows that emanated from the cone, including pressure ridges and hexagonal cooling joints. This flows and the cinder cone look amazingly fresh. The lava flows have been dated at between 6000 and 500 years old.

InsideAmboy031213
Verbena031213The warm temperatures at Amboy brought out some reptiles, to our delight. We saw at least three desert iguanas and many flowers, including the Sand Verbena (Albronia villosa) shown above. It felt like summer, although we were still in the middle of March. Tomorrow is expected to be even warmer. (Yes!)

Wooster Geologists in ancient lakes

March 11th, 2013

Manix031113ZZYZX, CALIFORNIA–Today Team Mojave studied the remains of ancient lake systems in the dry, dry desert. Early in the morning we drove west from Zzyzx to the Harvard Road exit off Interstate 15 and took a series of sandy roads to Bulwada Ridge on the shoreline of ancient Lake Manix. This Pleistocene, pluvial lake occupied a huge basin east of Barstow, California. Above we see laminated muds deposited in the lake itself. In the foreground are bouldery shoreline sediments. In the background you can see a scarp cut by the modern Mojave River (all underground at this point). Lake Manix was fed by the ancient Mojave River and supported a diverse fauna of fish and invertebrates.

MeagenBeachRidge031113Just to the north of the shoreline are long, twisting gravel bodies. Meagen Pollock is pondering one which has been truncated by modern erosion and now is topped by a beautiful desert pavement. (She found a beautifully polished artifact on one of these surfaces, by the way. It is a scraper with one bifacially-worked edge.) These gravel deposits are ancient beach ridges made by storm waves on Lake Manix.

OwlCanyonGroup031113After our visit to Lake Manix, we traveled to much older lake and river deposits in the Barstow Formation (Upper Miocene) exposed in Owl Canyon near Barstow. Our tradition here is that the students first explore the exposures and then bring back rocks to the picnic area for group analysis and discussion. The students above are clearly happy with all that they learned. (The two distinguished people in the background, by the way, are my parents Gary and Corinne Wilson who came to enjoy the geological fellowship.)

OwlCanyonTeaching585

A view from the other side of the table. Photo by Gary Wilson.

DesiccationCracks031113Kyle Burden found this beautiful example of two-stage desiccation cracks, preserved here as molds in the overlying sandstone.

RainbowBasin031113Another Wooster tradition is the group photograph in front of the outstanding syncline in the Barstow Formation exposed in nearby Rainbow Basin.

AftonSide031113Our final stop was in Afton Canyon about 50 miles east of Barstow. This narrow passageway is where Lake Manix drained into Soda and Silver Lakes, apparently more than once in a catastrophic manner. We crossed the mighty Mojave Rover on foot and then walked up this side canyon to check out the flood deposits and the lake deposits high on the skyline. These mark the highest level Lake Manix reached before it overtopped its dam and drained very quickly.

Another stimulating day which ended with a chicken dinner at the Desert Studies Center. Tomorrow we will return to the Barstow region but this time concentrating on structural issues and volcanic and plutonic rocks.

 

Death Valley Days

March 10th, 2013

BadwaterGroup031013DEATH VALLEY, CALIFORNIA–All geologists love Death Valley. No other place on Earth has such extraordinarily diverse geology combined with a modern infrastructure and a century of scientific study. The Wooster Geologists had a spectacular time in and around the valley today. Here we are above with the traditional group shot at Badwater. The weather could not have been better.

Zabriskie031013We left the Desert Studies Center at Zzyzx just after breakfast and drove through Baker and Shoshone to the southern end of Death Valley, seeing many wonderful sites. After lunch at the new National Park Service Furnace Creek Visitor Center, we then drove east and up out of the valley to Zabriskie Point. The above view has been published countless times by geologists and nature enthusiasts, but it has not lost its graphic power. We are looking here to the west at deeply eroded lake sediments of the Furnace Creek Formation. Towards the back of the light-colored material you can just make out the black streak of a basaltic intrusion.

ZabriskieOtherSide031013I think the other side of Zabriskie Point — the side looking out over Death Valley — is even more impressive. We see again the Furnace Creek Formation lake sediments, this time with alluvial deposits on top (visible on the right). These materials accumulated in an ancient lake and were lifted up and tilted by the tremendous faulting that formed Death Valley. The pinnacle is called Manly Beacon.

ZabriskieStudents031013

We took advantage of the sunlight and high spirits to take a picture of our Desert Geology 2013 students.

DanteView031013

We continued east and then south to Dante’s View, where we looked down into Death Valley from the dizzy heights. In this image we see Telescope Peak towering at 11,049 feet of elevation, while much of the valley floor below is lower than sealevel.

DanteViewFan031013Looking straight down from Dante’s View to Badwater (on the far right), we can see a complete alluvial fan from the narrow channel in the mountain slope to the spreading apron of debris over the salt pan on the valley floor. Badwater Road skirts the periphery of the fan.

After Dante’s View, we continued east and returned to Zzyzx via Death Valley Junction, Shoshone and Baker. Again, I can think of nowhere else one can see so much geological diversity in a single day, from the steamy floor of Death Valley to the heights above where we could walk through patches of snow.

Wooster’s Fossil of the Week: A brittle star from the Upper Jurassic of Germany

March 10th, 2013

Ophiopetra lithographica aboral larger 010813_585Wooster geologists have again greatly benefited from the donation of a collection by an alumnus. George Chambers (’79), a successful professional photographer, sent us several boxes of minerals, rocks and fossils he had acquired in his lifelong passion for geology. (George was a geology major at Wooster in the class just after mine.) Among the many world-class specimens he gave us are two fossil ophiuroids (brittle stars). They are Ophiopetra lithographica Enay and Hess, 1962, from the Lower Hienheim Beds (Lower Tithonian, Upper Jurassic) near Regensburg, Germany. They are part of the “Fossillagerstätte Hienheim“, a preserved brittle star ecosystem in a lagoon at the edge of a Late Jurassic sea. This is the same set of lithographic limestones in which the famous bird fossil Archaeopteryx was found.
Ophiopetra lithographica 010813_585In both these images you see the spiny arms of the brittle stars twisted about. It is their flexibility and snake-like movements in life that provoked the scientific name ophiuroids (serpent-forms) for the brittle stars. The “brittle” term comes from their ability to autotomize (spontaneously detach) their arms when threatened, leaving a squirming distraction for a predator as they escape.
Ophiopetra lithographica aboral 010813_585Ophiopetra lithographica is probably the most common fossil brittle star known. It was preserved by the countless millions in these Jurassic lagoons in Germany. Most geologists believe they were buried by fine-grained carbonate sediment suspended by sudden storms. As you can see in the above close-up, the preservation of the plates and spines is remarkable.

Most brittle stars are suspension feeders (sorting out food particles from the water), deposit feeders (eating organic material in the sediment) or scavengers. Ophiopetra lithographica may have been a carnivore with its heavily-spined arms and strong jaws. It likely ate small arthropods on the seafloor.

The evolution of brittle stars is interesting and controversial. They were relatively common in the Paleozoic and then just barely survived the Permian extinctions. Their rapid evolution into a variety of taxa in the Mesozoic and Cenozoic has led to many debates about their phylogeny. Even the placement of Ophiopetra into a family is a problem. Does it belong to the Family Aplocomidae where it was originally placed or to the older Family Ophiolepididae as has been recently suggested?

Our students will enjoy these fine fossils in the invertebrate paleontology course. They have doubled our collection of brittle stars! Thank you again to George Chambers for his thoughtfulness and generosity.

References:

Enay, R. and Hess, H. 1962. Sur la découvertes d’Ophiures (Ophiopetra lithographica n.g. n.sp.) dans le Jurassique supérieur du Haut-Valromey (Jura méridional). Eclogae geologicae Helvetiae 55: 657-678.

Hess, H. and Meyer, C.A. 2008. A new ophiuroid (Geocoma schoentalensis sp. nov.) from the Middle Jurassic of northeastern Switzerland and remarks on the Family Aplocomidae Hess 1965. Swiss Journal of Geosciences 101: 29-40.

Röper, M. and Rothgänger, M. 1998. Die Plattenkalke von Hienheim (Landkreis Kelheim) – Echinodermen-Biotope im Südfränkischen Jura. Eichendorf (Eichendorf Verlag), 110 S.

Stöhr, S. 2012. Ophiuroid (Echinodermata) systematics—where do we come from, where do we stand and where should we go? In: Kroh, A. and Reich, M. (Eds.) Echinoderm Research 2010: Proceedings of the Seventh European Conference on Echinoderms, Göttingen, Germany, 2–9 October 2010. Zoosymposia, 7: 147-161.

Thuy, B., Klompmaker, A.A. and Jagt, J.W.M. 2012. Late Triassic (Rhaetian) ophiuroids from Winterswijk, the Netherlands; with comments on the systematic position of Aplocoma (Echinodermata, Ophiolepididae). In: Kroh, A. and Reich, M. (Eds.) Echinoderm Research 2010: Proceedings of the Seventh European Conference on Echinoderms, Göttingen, Germany, 2–9 October 2010. Zoosymposia, 7: 163-172.

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