Wooster Paleontologists return to Caesar Creek Lake

September 6th, 2015

1 Shoreside collecting 090615Ohio is a wonderful place for paleontologists. One of the reasons is the thick, productive set of Upper Ordovician rocks that are exposed in the southwest of the state in and around Cincinnati. It is an easy drive south from Wooster into some of the most fossiliferous sediments in the world. Today Wooster’s Invertebrate Paleontology class went to Caesar Creek Lake in Warren County with its shoreline and emergency spillway exposing richly productive limestones and shales of the Waynesville, Liberty and Whitewater Formations (all of which equal the Bull Fork Formation). I’ve been there many times with many classes. The weather today was hot and dry — a contrast with last year’s torrential rains and mud.

2 Caesar Creek Lake Visitor Center MS 090615After a three-hour drive, we stopped first at the U.S. Army Corps of Engineers visitor center for Caesar Creek Lake. The Corps built the dam and spillway for the lake, and continues to maintain them both. This center has a set of museum displays and is where we obtain our fossil collecting permit. (This image was taken by Mara Sheban, a sophomore at The College of Wooster who will be contributing photographs to this blog and other geology projects.)

3 Initial briefing MS 090615Our initial briefing, with a review of the local geology and most common fossils. Since this is the start of the Invertebrate Paleontology course, most students have only begun to explore the topic. The fossils they collect on this trip will be the basis of a semester-long project of systematics and paleoecological analysis. (Photo by Mara Sheban.)

4 Marching to the outcrop 090615Our march to the far north of the exposures at the Caesar Creek Lake emergency spillway. We actually drop down from the spillway to the lake’s south shore. (See photo at the top of this post.) In the distant parking lot you can just make out a white pick-up truck in which the seasonal ranger is keeping an eye on us.

5 Jacob Nowell collecting MS 090615Jacob Nowell collecting small fossils washed from the shale along the lakeshore. (Photo by Mara Sheban.)

6 Jacob Pries trilobite MS 090615Jacob Pries found a nice enrolled Flexicalymene trilobite. Caesar Creek has a reputation as being a good place to find trilobites. We love them, but are also interested in the rest of the fossil fauna. (Mara Sheban image.)

7 Brachiopod hash 090615Some of the limestone units are a nearly solid hash of cemented brachiopod shells.

8 Hardground full view 0090616My favorite slabs at Caesar Creek Lake are the abundant carbonate hardgrounds like the above. A hardground is a cemented seafloor, usually with borings and encrusting organisms. This one began as a burrowed soft carbonate sediment. The burrows were filled with fine mud that cemented early on the seafloor. The surrounding softer matrix washed away, leaving a hardground with the burrows now in positive relief. Brachiopods and corals then lived in the nooks and crannies of this hard rock on the bottom of the sea.

9 Hardground closer 090615Here is the burrow system in a closer view. In the upper right is a beautiful encrusted rugose coral, shown in detail below.

10 Bryo encrusted rugosan 090615That coral is almost completely covered by a trepostome bryozoan. I would have loved to collect this specimen for further study, but the slab is too large and no tools are allowed on this outcrop.

In the coming weeks we will identify the fossils we gathered, apply to them several paleontological techniques such as cleaning, cutting, polishing and photography, and then put together a grand paleoecological analysis. We will be greatly assisted by two fantastic websites, one by Alycia Stigall at Ohio University called The Digital Atlas of Ordovician Life, and the other by Steve Holland at the University of Georgia titled The Stratigraphy and Fossils of the Upper Ordovician near Cincinnati, Ohio.

Wooster’s Fossil of the Week: Small and common orthid brachiopods from the Upper Ordovician of Ohio

August 7th, 2015

Cincinnetina meeki (Miller, 1875) slab 1 585
One of the many benefits of posting a “Fossil of the Week” is that I learn a lot while researching the highlighted specimens. I not only learn new things, I learn that some things I thought I knew must be, shall we say, updated. The above slab contains dozens of brachiopods (and a few crinoid ossicles and bryozoans). I have long called the common brachiopod here Onniella meeki. Now I learn from my colleagues Alycia Stigall and Steve Holland at their great Cincinnatian websites that since 2012 I should be referring to this species as Cincinnetina meeki (Miller, 1875). Jisuo Jin sorted out its taxonomy in a Palaeontology article three years ago:

Phylum: Brachiopoda
Class: Rhynchonellata
Order: Orthida
Family: Dalmanellidae
Genus: Cincinnetina
Species: Cincinnetina meeki (Miller, 1875)
Cincinnetina meeki (Miller, 1875) slab 2 585This slab, which resides in our Geology 200 teaching collection, was found at the famous Caesar Creek locality in southern Ohio. It is from the Waynesville/Bull Fork Formation and Richmondian (Late Ordovician) in age.
Cincinnetina meeki (Miller, 1875) slab 3 585You may see some bryozoans in this closer view. This bed is a typical storm deposit in the Cincinnatian Group. The shells were tossed about, most landing in current-stable conditions, and finer sediments were mostly washed away, leaving this skeletal lag.


Jin, J. 2012. Cincinnetina, a new Late Ordovician dalmanellid brachiopod from the Cincinnati type area, USA: implications for the evolution and palaeogeography of the epicontinental fauna of Laurentia. Palaeontology 55: 205–228.

Wooster’s Fossils of the Week: An Upper Ordovician cave-dwelling bryozoan fauna and its exposed equivalents

July 3rd, 2015

1 Downwards 063015This week’s fossils were the subject of a presentation at the 2015 Larwood Symposium of the International Bryozoology Association in Thurso, Scotland, last month. Caroline Buttler, Head of Palaeontology at the National Museum Wales, Cardiff, brilliantly gave our talk describing cryptic-and-exposed trepostome bryozoans and their friends in an Upper Ordovician assemblage I found years ago in northern Kentucky. They were the subject of an earlier Fossil of the Week post, but Caroline did so much fine work with new thin sections and ideas that they deserve another shot at glory. We are now working on a paper about these bryozoans and their borings. Below you will find the abstract of the talk and a few key slides to tell the story.


Trepostome bryozoans have been found as part of an ancient cave fauna in rocks of the Upper Ordovician (Caradoc) Corryville Formation exposed near Washington, Mason County, Kentucky.

Bryozoans are recognized as growing from the ceiling of the cave and also from an exposed hardground surface above the cave. Multiple colonies are found overgrowing one another and the majority are identified as Stigmatella personata. Differences between those growing upwards and those growing down from the roof have been detected in the limited samples.

The colonies have been extensively bored, these borings are straight and cylindrical. They are identified as Trypanites and two types are recognised. A smaller variety is confined within one colony overgrowth and infilled with micrite. In thin section it is observed that the borings follow the lines of autozooecial walls and do not cut across. This creates a polygonal sided boring, suggesting that the colonies were not filled with calcite at the time of the boring. The second variety has a larger tube size and its infilling sediment has numerous dolomite rhombs and some larger fossil fragments including cryptostomes, shell and echinoderm pieces. These cut through several layers of overgrowing bryozoans. Some of the borings contain cylindrical tubes of calcite similar to the ‘ghosts’ of organic material described by Wyse Jackson & Key (2007).

Very localised changes in direction of colony growth due to an environmental effect are seen.

Bioclaustration in these samples provides evidence for fouling of the colony surface, indicating that the bryozoans overgrew unknown soft-bodied organisms.


Wyse Jackson, P. N., and M. M. Key, Jr. (2007). Borings in trepostome bryozoans from the Ordovician of Estonia: two ichnogenera produced by a single maker, a case of host morphology control. Lethaia. 40: 237-252.

2 Title 0630153 Location 0630154 Strat position 0630155 hdgd up 0630156 hdgd down 0630157 Growth up 0630158 Growth down 0630159 Stigmatella 06301510 Cartoon 06301511 Boring A 06301512 Boring B 06301513 Ghosts explanation14 Ghosts 06301515 Overgrowths 06301516 Further questions 063015

Wooster’s Fossils of the Week: An encrusted bivalve external mold from the Upper Ordovician of Indiana

June 26th, 2015

1 Anomalodonta gigantea Waynesville Franklin Co IN 585I love this kind of fossil, which explains why you’ve seen so many examples on this blog. We are looking at an encrusted external mold of the bivalve Anomalodonta gigantea found in the Waynesville Formation exposed in Franklin County, Indiana. I collected it many years ago as part of an ongoing study of this kind of preservation and encrustation.
2 Anomalodonta gigantea Waynesville Franklin Co IN 585 annotatedTo tell this story, I’ve lettered the primary interest areas on image above. First, an external mold is an impression of the exterior of an organism. In this case we have a triangular clam with radiating ribs in its shell. The exterior of the shell with its ribs was buried in sediment and the shell dissolved, leaving the basic impression above. It is a negative relief. Please now refer to the letters for the close-up images below.

3 Bryo Anomalodonta gigantea Waynesville Franklin Co INA. At the distal end of the bivalve mold is what looks at first to be the original shell. It is calcitic, though, and we know this bivalve had an aragonitic shell. A closer look shows that this is actually the attaching surface of an encrusting bryozoan that bioimmured the original bivalve shell, which has since dissolved away. This smooth surface is the bryozoan underside; we see the characteristic zooecia (tubes holding the individual zooids) only when this surface is weathered away.

4 Borings Anomalodonta gigantea Waynesville Franklin Co INB. These tubular objects are infillings of borings (maybe Trypanites)that were cut into the original aragonitic shell of the bivalve. The tunnels of the borings were filled with fine sediment, and then the shell dissolved away, leaving these casts of the borings.

5 Inarticulate scar Anomalodonta gigantea Waynesville Franklin Co INC and D. In the middle of the external mold is this curious circular feature (C) mostly surrounded by a bryozoan (D). There was at one time a circular encruster, likely an inarticulate brachiopod like Petrocrania, that sat directly on the external mold surface. The bryozoan colony grew around but not over it because it was alive and still opening and closing its valves for feeding. The bryozoan built a vertical sheet of skeleton around it as a kind of sanitary wall. You may be able to see the other three or four structures in the top image showing brachiopod encrusters that left the building. This is an example of fossils showing us a living relationship, even if one is not longer preserved.

This fossil and its sclerobionts (hard substrate dwellers) show us that soon after the bivalve died its aragonitic shell dissolved away, leaving as evidence the external mold in the sediment, the bioimmuring bryozoan, and the boring casts. Very soon thereafter bryozoans and brachiopods encrusted the available hard substrate. This is a typical example of early aragonite dissolution on the sea floor during a Calcite Sea interval.


Palmer, T.J. and Wilson, M.A. 2004. Calcite precipitation and dissolution of biogenic aragonite in shallow Ordovician calcite seas. Lethaia 37: 417-427.

Taylor, P.D. 1990. Preservation of soft-bodied and other organisms by bioimmuration—a review. Palaeontology 33: 1-17.

Taylor, P.D. and Wilson, M.A. 2003. Palaeoecology and evolution of marine hard substrate communities. Earth-Science Reviews 62: 1-103.

Wilson, M.A., Palmer, T.J. and Taylor, P.D. 1994. Earliest preservation of soft-bodied fossils by epibiont bioimmuration: Upper Ordovician of Kentucky. Lethaia 27: 269-270.

Wooster’s Fossil of the Week: An undescribed cyclostome bryozoan from the Upper Ordovician of Oklahoma

June 19th, 2015

HT_1276 585Paul Taylor and I presented a talk this month at the Larwood Symposium of the International Bryozoology Association in Thurso, Scotland. (Yes, way in the tippy-top of Scotland. Very cool.) Paul found the above wiggly bryozoan encrusting the interior of an orthid brachiopod identified as Multicostella sulcata (thanks, Alycia Stigall!) in the Lower Echinoderm Zone of the Mountain Lake Member of the Bromide Formation (Upper Ordovician, Sandbian) near Fittstown, Oklahoma. This bryozoan is “new to science”, as we grandly say. Paul generously invited me to describe it with him in this presentation and in a future paper. We did a 1994 paper together on Corynotrypa, a similar cyclostome bryozoan. The following are a few slides from our Larwood talk.






Slide21_052815This last image showing what appear to be an interior wall with a pore is critical. Corynotrypa does not have such walls, so our bryozoan is more like a sagenellid cyclostome.


Carlucci, J.R., Westrop, S.R., Brett, C.E. and Burkhalter, R. 2014. Facies architecture and sequence stratigraphy of the Ordovician Bromide Formation (Oklahoma): a new perspective on a mixed carbonate-siliciclastic ramp. Facies 60: 987-1012.

Taylor, P.D. and Wilson, M.A. 1994. Corynotrypa from the Ordovician of North America: colony growth in a primitive stenolaemate bryozoan. Journal of Paleontology 68: 241-257.

Wooster’s Fossil of the Week: A bored and formerly encrusting trepostome bryozoan from the Upper Ordovician of Indiana

March 20th, 2015

1 Trep Upper 030115The lump above looks like your average trepostome bryozoan from the Upper Ordovician. I collected it from the Whitewater Formation of the Cincinnatian Group at one of my favorite collecting sites near Richmond, Indiana. In this view you can just barely make out the tiny, regular holes that are the zooecia (calcitic tubes that held the bryozoan individuals — the zooids). There are bits of other fossils stuck to the outside, so it’s not particularly attractive as fossils go. (Except that all fossils are fascinating messengers in time.)

2 Trep Upper CloseWith this closer view you can see my initial interest in this particular bryozoan. Again, the regular, tiny holes are the zooecia. The larger pits are borings by worm-like, filter-feeding organisms. These borings are either in the ichnogenus Trypanites (if they are cylindrical) or Palaeosabella (if they are clavate, meaning clubbed at their distal ends). Such borings are common in all types of skeletal fossils in the Upper Ordovician — so common that they are part of the evidence for the Ordovician Bioerosion Revolution. So, let’s flip this ordinary, bored bryozoan over and see what’s underneath:

3 Trep Under 030115Here’s the main scientific beauty! We’re looking at the underside of the bryozoan. Ordinarily we’d expect to see a shell here that the bryozoan was encrusting, but the shell is gone. We’re gazing directly at the attachment surface of the bryozoan. It’s as if the colony had encrusted a sheet of glass and we’re looking right through it. The shell it was originally attached to has been removed either through dissolution (it might have been an aragonitic bivalve) or physical removal (it may have been a calcitic brachiopod). The borings are now much more prominent. They penetrated through the bryozoan into the mysterious missing shelly substrate. Some are small pits that just intersected the shell, others are horizontal as the boring organism turned at a right angle when it reached the shell and drilled along the bryozoan-shell interface. Removing the shell exposed the distal parts of these borings — parts that ordinarily would have been hidden by the encrusted shell.

4 Trep Under labeledHere is a closer, labeled view of this bryozoan basal surface. A is the earliest encruster recorded in this scenario; it is a small encrusting bryozoan that was first on the shelly substrate and then completely overgrown (or bioimmured) by the large trepostome. B shows that the trepostome was growing on a shell that already had borings from a previous encruster-borings combination that must have fallen off; these are grooves in the substrate that the trepostome filled in as it covered the shell. C is one of the many later borings that cut perpendicularly through the bryozoan and worked along the shell-bryozoan interface; as described above, only when that shelly substrate was removed would these be visible. In this surprisingly complex story, B represents an earlier version of C. We thus know that the shell was encrusted by one bryozoan, bored, and then that bryozoan was freed at its attachment (and not found in our collection). The same shell was then encrusted by this second bryozoan, which recorded the groove (or “half-borings”) made during the first encrustation.

These half-borings were first described in 2006 when my students Cordy Dennison-Budak and Jeff Bowen worked with me on them and we had a GSA abstract. Coleman Fitch is presently completing his Senior Independent Study enlarging the database for these features and developing detailed interpretations. The main implication from this work is that thick trepostome bryozoan encrusters often “popped off” shells, leaving no signs of their presence unless there were these half-borings in the shell surfaces and bryozoan undersides. Paleoecology and taphonomy on a very small scale!


Taylor, P.D. 1990. Preservation of soft-bodied and other organisms by bioimmuration—a review. Palaeontology 33: 1-17.

Wilson, M.A., Dennison-Budak, W.C., and Bowen, J.C. 2006. Half-borings and missing encrusters on brachiopods in the Upper Ordovician: Implications for the paleoecological analysis of sclerobionts. Geological Society of America Abstracts with Programs, Vol. 38, No. 7, p. 514.

Wilson, M.A. and Palmer, T.J. 2006. Patterns and processes in the Ordovician Bioerosion Revolution. Ichnos 13: 109-112.

Wilson, M.A., Palmer, T.J. and Taylor, P.D. 1994. Earliest preservation of soft-bodied fossils by epibiont bioimmuration: Upper Ordovician of Kentucky. Lethaia 27: 269-270.


Wooster’s Fossils of the Week: Beautiful trace fossils from the Upper Ordovician of southern Ohio

December 19th, 2014

Trace fossils Bull Fork Ordovician OH 585Every year we highlight at least one of the fossils found and studied by Wooster’s Invertebrate Paleontology class as part of their field and laboratory exercises. This year it is this nice slab of trace fossils collected by Curtis Davies (’15) on our August 31 field trip to the emergency spillway in Caesar Creek State Park. I didn’t even notice it at the time Curtis picked it up. I only saw its full glory when he photographed the rock as part of a paleontological essay.
CurtisGalen083114aCurtis Davies is the smiling, bearded guy in the back (with Galen Schwartzberg) at the Caesar Creek outcrop. The rain had finally stopped and everyone was happy.

The traces are exposed here on the bottom of a bed of argillaceous limestone. They are preserved in what trace fossil workers (ichnologists) call convex hyporelief, which means simply that they stick out on the base (or sole) of the rock slab. These were tunnels originally excavated in soft mud by worm-like animals. The tunnels were filled with sediment that cemented up more resistant than the surrounding matrix, and thus were weathered in this relief.
Taenidium serpentinum Heer, 1877Most of the trace fossils here are the simple unlined burrow called Planolites, one of the most common traces in the Ordovician of the Cincinnati area. The trace labelled with the red “T” above, though, is rare here. Note that it is formed by a series of pulse-like movements that produced segments in the sediment infill. My estimate is that this trace can be classified as Taenidium serpentinum Heer, 1877. It is not common in the Ordovician.
Heer, Oswald, 1809-1883Oswald Heer (1809-1883), the scientist who named Taenidium serpentinum, was a Swiss geologist and botanist. As was the case for many educated Europeans, he started as a clergyman, even signing up for holy orders. The natural world captivated him, though, and starting with insects he worked his way up to become a naturalist and professor of botany at the University of Zürich. He was one of the key figures in the establishment of paleobotany (the study of fossil plants).
Taenidium serpentinum Heer, 1877 image 585Here is Heer’s figure of Taenidium serpentinum from Plate XLV in his 1877 book, Flora fossilis Helvetiae (Fossils Plants of Switzerland). You see the irony already. Heer described this trace fossil as a plant, inadvertently becoming one of the early figures in ichnology, the study of trace fossils.

Oswald Heer published many books and papers, becoming well known for his geological and paleontological explorations and descriptions. He was awarded the prestigious Wollaston Medal from the Geological Society of London in 1874. He was an earlier advocate of using fossils to sort on problems of paleogeography. He knew, for example, that Miocene fossils in Europe and North America were very similar, so he suggested in those days before Plate Tectonic Theory that the two continents were connected by a “land bridge“. This was called the “Atlantis Hypothesis”, and you can imagine the confusion that name caused among various cranks and pseudoscientists looking for Plato’s mythical continent. Heer died in Switzerland in 1883.


D’Alessandro, A. and Bromley, R.G. 1987. Meniscate trace fossils and the Muensteria-Taenidium problem. Palaeontology 30: 743-763.

Heer, O. 1877. Flora fossilis Helvetiae: Die vorweltliche flora der Schweiz. Zürich, J. Wurster & Company. 182 p.

Keighley, D.G. and Pickerill, R.K. 1994. The ichnogenus Beaconites and its distinction from Ancorichnus and Taenidium. Palaeontology 37: 305-338.

Keighley, D.G. and Pickerill, R.K. 1995. Commentary: The ichnotaxa Palaeophycus and Planolites: Historical perspectives and recommendations. Ichnos 3: 301-309.

Wooster’s Fossils of the Week: A trace fossil from the Ordovician of Estonia

November 21st, 2014

Hyoliths03_585The fossils above have been in a previous post as examples of hyolith internal molds from the Middle Ordovician of northern Estonia. I collected them on my first visit to the Baltic countries in 2006. This week I want to recognize them again, but this time for the squiggly trace fossils you can just make out on their outer surfaces. These are the ichnospecies Arachnostega gastrochaenae Bertling, 1992. They are the subject of a paper that has just appeared in Palaeontologia Electronica entitled, simply enough, “The trace fossil Arachnostega in the Ordovician of Estonia (Baltica)“. The senior author is my Estonian buddy Olev Vinn. My Polish friend Michał Zatoń, my new Estonian colleague Ursula Toom, and I are co-authors.
399-861 copyAbove is an unpublished image of a gastropod internal mold from the Estonian Ordovician taken by Olev. It shows very well the variable branching nature Arachnostega. This trace was formed by a deposit-feeding organism mining organic material in a sediment-filled shell. It worked along the sediment-shell interface, probably because there was more nutrient value at that margin. The internal mold was formed when sediment filling the shell was cemented and the shell dissolved away, leaving the hard mold behind.
Screen Shot 2014-11-02 at 4.05.40 PMThis is Figure 3.1 in the new paper. Note the variation in the traces as well as the shells it inhabited. The caption as published: Arachnostega gastrochaenae Bertling in a gastropod from Haljala Regional Stage (Sandbian), Aluvere Quarry, northern Estonia. GIT 399-948-1. 2. Arachnostega gastrochaenae Bertling in a gastropod from the Kunda Regional Stage (Darriwilian), Kunda Ojaküla, northern Estonia. GIT 404-355-1. 3. Arachnostega gastrochaenae Bertling in a bivalve from the Haljala Regional Stage (Sandbian), Aluvere Quarry, northern Estonia. GIT 399-1590-1. 4. Arachnostega gastrochaenae Bertling in a bivalve from the Haljala Regional Stage (Sandbian), Aluvere Quarry, northern Estonia. GIT 399-1601-1. 5. Arachnostega gastrochaenae Bertling in a cephalopod from the Uhaku Regional Stage (Darriwilian), Püssi, northern Estonia. GIT 695-12-1.

Our paper analyzes the distribution of Arachnostega through the Ordovician of Baltica, a paleocontinent with a long history, including a collision with Avalonia (western Europe today, more or less) in the Late Ordovician. By plotting the occurrences of Arachnostega over time, we conclude that the makers of Arachnostega likely preferred cool climates and bivalve shells over gastropods. The tracemakers may have also been negatively influenced by the many biotic changes associated with the Great Ordovician Biodiversification Event.

Please check out the article itself. As with all papers in Palaeontologia Electronica, it is open access.


Bertling, M. 1992. Arachnostega n. ichnog. – burrowing traces in internal moulds of boring bivalves (late Jurassic, northern Germany). Paläontologische Zeitschrift 66: 177-185.

Vinn, O., Wilson, M.A., Zatoń, M. and Toom, U. 2014. The trace fossil Arachnostega in the Ordovician of Estonia (Baltica). Palaeontologia Electronica 17, Issue 3; 41A; 9 p.

Wooster’s Fossil of the Week: Upper Ordovician bivalve bioimmured by a bryozoan

November 7th, 2014

DSC_4503This week’s fossil is a simple and common form in the Cincinnatian Series (Upper Ordovician) of the Ohio, Indiana and Kentucky tri-state area. We are looking above at the base of a trepostome bryozoan that encrusted the outside of an aragonite bivalve shell. The bivalve shell (probably a species of Ambonychia) dissolved away, leaving its impression in the base of the calcitic bryozoan. This fossil is from the Upper Whitewater Formation (Richmondian) in eastern Indiana near Richmond itself.
DSC_4516In this closer view you can see the plications (“ribs”) of the bivalve preserved in negative relief on the attachment surface of the bryozoan. Close examination shows the individual zooecia of the bryozoan exquisitely molding the bivalve topography.

This is a kind of substrate bioimmuration, a preservational mode in which a skeletal organism (the bryozoan here) overgrows another organism (with a soft body or hard skeleton), making an impression of it in its base. The overgrown organisms is rots or dissolves away, leaving the exposed mold. You can also think of it as a kind of external mold produced by a living organism (the encruster). Such “vital immuration” was first described by Vialov (1961), and it is thoroughly covered by Paul Taylor in his 1990 paper cited below.

Again, these fossils are common in the Cincinnatian, and this one is far from being the fanciest. It is the Fossil of the Week because of its very ordinary nature, yet it provides extraordinary information. The aragonitic shell the bryozoan encrusted would have been lost forever after it dissolved if this bryozoan hadn’t occupied it and built a calcitic memorial. I’ve collected now hundreds of these substrate bioimmurations, and they have been critical in many studies, from the preservation of soft-bodied sclerobionts (see Wilson et al., 1994) to the revelation of boring interiors (and thus the behavior of the borers) and skeletal sclerobiont paleoecology. I’m also convinced there are many aragonitic mollusk taxa in the Cincinnatian that are known only through this bioimmuration process. These are fascinating fossils my students and I will continue to collect and study.


Taylor, P.D. 1990. Preservation of soft-bodied and other organisms by bioimmuration—a review. Palaeontology 33: 1-17.

Vialov, O.S. 1961. Phenomena of vital immuration in nature. Dopovidi Akademi Nauk Ukrayin’ skoi RSR 11: 1510-1512.

Wilson, M.A., Palmer, T.J. and Taylor, P.D. 1994. Earliest preservation of soft-bodied fossils by epibiont bioimmuration: Upper Ordovician of Kentucky. Lethaia 27: 269-270.

Wooster’s Fossils of the Week: A pair of molded nautiloids from the Upper Ordovician of northern Kentucky

October 24th, 2014

1 Nautiloid pair 091314Two nautiloids are preserved in the above image of a slab from the Upper Ordovician of northern Kentucky. (I wish I knew which specific locality. This is why paleontologists are such fanatics about labeling specimens.) The top internal mold (meaning it is sediment that infilled a shell now dissolved away) has been covered in a previous blog entry. This week I want to concentrate on the nautiloid at the bottom.

These nautiloids belong to the Family Orthoceratidae McCoy, 1844, which existed from the Early Ordovician (490 million years ago) through the Triassic (230 million years ago). They had conical, aragonitic shells with walls inside separating chambers (camerae) and a central tube (the siphuncle) connecting them. They were swimming (nektic) predators that could control their buoyancy through a mix of gases and liquids in the camerae mediated by the siphuncle.

What is most interesting here is the preservation of these nautiloids. The aragonitic shells were dissolved away at about the same time the internal sediment was cemented, forming the internal molds. These molds were exposed on the seafloor, attracting encrusting organisms. This means the dissolution and cementation took place quickly and in the marine environment, not after burial. This rapid dissolving of aragonite and cementation by calcite is typical of Calcite Sea geochemistry, something we don’t see in today’s Aragonite Seas.
2 Nautiloid siphuncle 091314Above is a close view of the cemented siphuncle of the lower nautiloid, heavily encrusted by a trepostome bryozoan.
3 Bryozoan undersideEven more cool, the outside of the lower nautiloid was encrusted by several trepostome bryozoan colonies. When the shell dissolved it left the undersides of these bryozoans exposed, as seen above. These undersides often contain the remains of shelly organisms the bryozoans encrusted (see the Independent Study project by Kit Price ’13) and even soft-bodied animals (epibiont bioimmuration; see Wilson et al., 1994).

A neat package here resulting from biological, sedimentological and geochemical factors.


Palmer, T.J., Hudson, J.D. and Wilson, M.A. 1988. Palaeoecological evidence for early aragonite dissolution in ancient calcite seas. Nature 335 (6193): 809–810.

Sweet, W.C. 1964. Nautiloidea — Orthocerida, in Treatise on Invertebrate Paleontology. Part K. Mollusca 3, Geological Society of America, and University of Kansas Press, New York, New York and Lawrence, Kansas.

Wilson, M.A., Palmer, T.J. and Taylor, P.D. 1994. Earliest preservation of soft-bodied fossils by epibiont bioimmuration: Upper Ordovician of Kentucky. Lethaia 27: 269-270.

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