Wooster’s Fossil of the Week: A bitten brachiopod (Upper Ordovician of southeastern Indiana)

February 5th, 2016

1 Best bitten Glyptorthis insculpta (Hall, 1847)This brachiopod, identified as Glyptorthis insculpta (Hall, 1847), was shared with me by its collector, Diane from New York State. She found it in a muddy horizon of the Bull Fork Formation (Upper Ordovician) in southeastern Indiana. She immediately noted the distorted plicae (radiating ribs) on the left side of this dorsal valve, along with the invagination along the corresponding margin. (Thanks for showing this to me, Diane, and allowing me to include it in this blog.)
2 Best closer Glyptorthis insculpta (Hall, 1847)Above  is a closer view of the unusual plicae. Note that they radiate from the top center of the brachiopod, extending as the shell grew outward along its margins. Something happened, though, when the brachiopod was growing. The shell was seriously damaged by a puncturing object. The brachiopod repaired the hole by closing it up with additional shell material coming from either side. The inwardly-curved plicae show the pattern of shell regrowth.
3 Reverse of best Glyptorthis insculpta (Hall, 1847)This is a view of the same brachiopod from the other side, showing that the ventral valve was damaged in the same event, but with slightly less destruction.

So how did such damage occur on that Ordovician seafloor? Some predator likely took a bite out of the brachiopod as it lay in its living position with the valves extended upwards into the seawater. Most brachiopods do not survive such events, but this one did.

Who was the probable predator? For that we turn to the work of the late Richard Alexander (1946-2006). He did the definitive study of pre mortem damage to brachiopods in the Cincinnatian Group in 1986, concluding that the most likely predators on these brachiopods were nautiloid cephalopods. Some of this figures show nearly identical healed scars on similar orthid brachiopods.
4. Richard AlexanderRichard Alexander was an accomplished paleontologist who lost his life in a swimming accident off the coast of St. Lucia just over nine years ago. He was born in Covington, Kentucky, right across the river from Cincinnati. As is so common with children in that part of the world, he developed a passion for fossils. He attended the University of Cincinnati, majoring in geology, He then went to Indiana University, completing a PhD dissertation titled: “Autecological Studies of the Brachiopod Rafinesquina (Upper Ordovician), the Bivalve Anadara (Pliocene), and the Echinoid Dendraster (Pliocene).” (We don’t see such diverse projects very much these days.) He taught at Utah State University from 1972 to 1980, and then at Rider University in New Jersey from 1981 until his death. He served as an administrator at several levels at Rider, and was known as an excellent teacher. His research interests changed when he moved to the East Coast, becoming increasingly focused on modern mollusks. No doubt he would still be contributing to paleontology but for the randomness of a freak wave in the Caribbean.


Alexander, R.R. 1981. Predation scars preserved in Chesterian brachiopods: probable culprits and evolutionary consequences for the articulates. Journal of Paleontology 55: 192-203.

Alexander, R.R. 1986. Resistance to and repair of shell breakage induced by durophages in Late Ordovician brachiopods. Journal of Paleontology 60: 273-285.

Dodd, J.R. 2008. Memorial to Richard Alexander (1946-2006). Geological Society of America Memorials 37: 5-7.

Wooster’s Fossil of the Week: A brachiopod with a heavy burden (Upper Ordovician of southeastern Indiana)

January 29th, 2016

1 Trepostome on Hebertella richmondensisYes, the above image doesn’t look much like a brachiopod, but just wait. We see a trepostome bryozoan with extended knobs and a few borings. Flip it over, though …
2 Hebertella richmondensis ventral view 585… and we see that the bryozoan almost entirely covers a brachiopod. So far, so common among Ordovician fossils. However, look closely at the margin of the brachiopod valve and how clearly it is delineated from the bryozoan. It is apparent that the bryozoan had encrusted a living brachiopod, and the brachiopod stayed alive, keeping the essential commissure (the gap between the valves) open for feeding. We are looking at the valve that was in contact with the substrate (the underside of the living brachiopod). The bryozoan occupied the upper exposed surface, growing across that valve (which is invisible to us now), past its edge, but not closing the gap with the other valve. The same bryozoan species is found on the above visible valve, but only as two thin films unconnected to the colony on the upper side.
3 Hebertella richmondensis bryo close annotatedA closer view of the brachiopod hinge shows additional evidence that the bryozoan and brachiopod were living together. The red arrow on the left points to where the fleshy pedicle (attaching stalk) of the brachiopod extended from the shell to meet the substrate. The bryozoan here curves around the now-vanished pedicle. The yellow arrow on the right shows how the bryozoan growth surface folded to accommodate the opening valves at the hinge. Pretty cool.

I can’t identify the bryozoan beyond Order Trepostomata without cutting it open. The brachiopod, though, appears to be Hebertella richmondensis Foerste, 1909. This specimen is from the Whitewater Formation (Upper Ordovician, upper Katian) exposed near Richmond, Indiana. It was collected on one of my field trips in 2003.
4 Hebertella richmondensis ventral view 585 annotatedWhat do we learn from this little assemblage? We first see a relatively uncommon example of a clear living relationship between a sclerobiont and its host. We also learn that the brachiopod could continue to open its valves for feeding despite the heavy calcitic bryozoan weighing it down. We even can see that this brachiopod was not living on a soft muddy substrate because only a small triangular-shaped area (see above) in the center was clear of encrusters; the thin bryozoan (and maybe a bit of the stromatoporid sponge Dermatostroma) had enough space between the valve and the substrate to feed and respire. None of this is surprising, but it is nice to see our models of how these organisms lived are congruent with the evidence.


Alexander, R.R. and Scharpf, C.D. 1990. Epizoans on Late Ordovician brachiopods from southeastern Indiana. Historical Biology 4: 179-202.

Foerste, A.F. 1909. Preliminary notes on Cincinnati fossils. Bulletin of the Scientific Laboratory of Denison University 14: 208–232.

Walker, L.G. 1982. The brachiopod genera Hebertella, Dalmanella, and Heterorthina from the Ordovician of Kentucky. USGS Professional Paper 1066-M.

Wright, D.F. and Stigall, A.L. 2013. Phylogenetic revision of the Late Ordovician orthid brachiopod genera Plaesiomys and Hebertella from Laurentia. Journal of Paleontology 87: 1107-1128.

Wooster’s Fossils of the Week: Atrypid brachiopods attached to a trepostome bryozoan from the Upper Ordovician of southern Indiana

January 8th, 2016

Zygospira Attached 585This is a follow-up post to our entry on Christmas Day two weeks ago. Above is a trepostome bryozoan (the long porous piece) with specimens of the atrypid brachiopod Zygospira modesta clustered around it. They are positioned with their ventral valves outward because in life they were attached to this bryozoan with tiny fleshy stalks called pedicles. They were buried quickly enough that this spatial relationship was preserved. Cool. This assemblage was found in the Liberty Formation (Upper Ordovician) exposed in a roadcut in southern Indiana.
Zygospira modesta dorsal annotatedThis is a view of the dorsal side of Zygospira modesta showing the pedicle opening in the ventral valve at the apex of the shell.


Copper, P. 1977. Zygospira and some related Ordovician and Silurian atrypoid brachiopods. Palaeontology 20: 295-335.

Sandy, M.R. 1996. Oldest record of peduncular attachment of brachiopods to crinoid stems, Upper Ordovician, Ohio, USA (Brachiopoda; Atrypida: Echinodermata; Crinoidea). Journal of Paleontology 70: 532-534.

Wooster’s Fossil of the Week: Tiny atrypid brachiopods from the Upper Ordovician of southern Ohio

December 25th, 2015

Zygospira modesta Waynesville 585These exquisite little brachiopods are among the most abundant fossils in the Upper Ordovician of the Cincinnati area. My Invertebrate Paleontology students collected dozens of them from the Waynesville Formation on our field trip to Caesar Creek Lake last semester. Their ubiquity, though, doesn’t make them any less precious.
Zygospira modesta dorsalThis is Zygospira modesta (Say in Hall, 1847). Above is a dorsal valve view of a single specimen. At the apex you can see a tiny round hole from which a fleshy pedicle extended to attach the brachiopod to a hard substrate.
Zygospira modesta ventralHere is the ventral valve view. Zygospira is an atrypid brachiopod, meaning that its internal support (brachidium) for the filter-feeding lophophore is looped in a characteristic way, shown below.
Hall diagram ZygospiraThe diagrams above are from Hall (1867) who named the genus Zygospira and wished to further distinguish it from other atrypid brachiopods.

The taxonomy of Zygospira modesta is a bit messy, as many early 19th Century species descriptions tended to be. It was apparently first named Producta modesta by Thomas Say (see below) but not actually published as such. James Hall described it as Atrypa modesta in 1847. Later in 1862 he named Zygospira as a new genus, making Z. modesta its type species but not indicating a type locality.
Thomas_Say_1818We met Thomas Say (1787-1834) earlier in this blog, recognizing him as the scientist who named Exogyra costata in 1820. He is shown above in an 1818 portrait. Say was a brilliant American natural historian. Among his many accomplishments in his short career, he helped found the Academy of Natural Sciences of Philadelphia in 1812, the oldest natural science research institution and museum in the New World. He is best known for his descriptive entomology in the new United States, becoming one of the country’s best known taxonomists. He was the zoologist on two famous expeditions led by Major Stephen Harriman Long. The first, in 1819-1820, was to the Great Plains and Rocky Mountains; the other (in 1823) was to the headwaters of the Mississippi. Along with his passion for insects, Say also studied mollusk shells, both recent and fossil. He was a bit of an ascetic, moving to the utopian socialist New Harmony Settlement in Indiana for the last eight years of his life. It is said his simple habits and refusal to earn money caused problems for his family. Say succumbed to what appeared to by typhoid fever when he was just 47.


Copper, P. 1977. Zygospira and some related Ordovician and Silurian atrypoid brachiopods. Palaeontology 20: 295-335.

Hall, J. 1862. Observations upon a new genus of Brachiopoda. Report New York State Museum, Natural History 15: 154-155.

Hall, J. 1867. Note upon the genus Zygospira and its relations to Atrypa. Report New York State Museum, Natural History 20: 267-268.

Sandy, M.R. 1996. Oldest record of peduncular attachment of brachiopods to crinoid stems, Upper Ordovician, Ohio, USA (Brachiopoda; Atrypida: Echinodermata; Crinoidea). Journal of Paleontology 70: 532-534.

Wooster’s Fossil of the Week: A tabulate coral from the Upper Ordovician of southern Ohio

December 18th, 2015

Calapoecia huronensis Billings, 1865 top 585We have here another fossil collected by a Wooster student on the August 2015 College of Wooster Invertebrate Paleontology field trip to Caesar Creek Lake, Ohio. Eduardo Luna picked up this specimen of the tabulate coral Calapoecia huronensis (Billings, 1865) from the Waynesville Formation (Upper Ordovician). For some reason in all my years of working in the Upper Ordovician, I’ve not come across this coral species before. Eduardo had sharp eyes as you can see it is rather small. The circular tubes are corallites, each of which held a coral polyp in life. This particular coral is distinctive for its septal spines along the inside rim of each corallite, giving them a beaded appearance.

Calapoecia huronensis Billings, 1865 bottom 585This is the underside of Eduardo’s coral. The corallites on the left side are eroded, showing the elongated septal spines that run lengthwise down their inside walls.

CNSPhoto-GEOLOGISTWe met the author of C. huronensis, Elkanah Billings (1820-1876), earlier this year, but why not show the handsome Canadian again? He originally described this coral species in 1865. He was Canada’s first government paleontologist, and he very much looked the part. Billings was born on a farm near Ottawa. He went to law school and became a lawyer in 1845, but he gave up stodgy law books for the bracing life of a field paleontologist. In 1856, Billings joined the Geological Survey of Canada, eventually naming over a thousand new species in his career. The Billings Medal is given annually by the Geological Association of Canada to the most outstanding of its paleontologists.


Billings, E. 1865. Notice of some new genera and species of Palaeozoic fossils. Canadian Naturalist and Geologist, New Series 2: 432–452.

Browne, R.G. 1965. Some Upper Cincinnatian (Ordovician) colonial corals of north-central Kentucky. Journal of Paleontology 39: 1177-1191.

Cox, I. 1936. Revision of the genus Calapoecia Billings. Bulletin of the National Museum of Canada 80: 1–48.

Jull, R.K. 1976. Review of some species of Favistina, Nyctopora, and Calapoecia (Ordovician corals from North America). Geological Magazine 113: 457-467.

Wooster’s Fossil of the Week: A common trilobite from the Upper Ordovician of Ohio

December 11th, 2015

Flexicalymene meeki cephalon view 585This beautiful specimen was collected by Wooster student Eve Caudill on this year’s College of Wooster Invertebrate Paleontology field trip to Caesar Creek Lake, Ohio. It is the iconic trilobite Flexicalymene meeki (Foerste, 1910) from a soft, “buttery” shale in the Waynesville Formation (Upper Ordovician). This is one of the most common trilobite species in the world, and it has been photographed thousands of times, so I posed it at an unconventional, rakish angle. We are looking here at the cephalon (head) of the animal. I like the way the remnants of the enclosing sedimentary matrix cling to the low places, highlighting the bumps and ridges. The center of the cephalon shows the distinctive glabella with its side lobes. The stomach of the trilobite was housed underneath it. The two eyes are visible on either side of the glabella, the one on the right split by the slightly-open facial suture used for dividing its exoskeleton during molting (ecdysis).

Flexicalymene meeki pygidium view 585This is a view of the pygidium (tail end) of the same Flexicalymene meeki specimen. It is tucked under the leading edge of the cephalon in the classic enrollment position. Trilobites likely enrolled for several reasons, but the primary one was almost certainly to affect a pill-bug-like defense against predators.

Flexicalymene meeki side view 585This is a side view of the enrolled trilobite. The articulated segments between the cephalon and pygidium constitute the thorax.foerste-1936We met the author of Flexicalymene meeki four years ago in this blog, so let’s visit him again. August F. Foerste (1862-1936) was one of the pioneers of Cincinnatian paleontology and stratigraphy. He grew up and worked in the Dayton, Ohio, area. Foerste went to Denison University where he was a very successful undergraduate, publishing several geological papers. He returned to Dayton after graduation with a PhD from Harvard, teaching high school for 38 years. When he retired he was offered a teaching position at the University of Chicago, but instead went to work at the Smithsonian Institution until the end of his life.

A final note from the Invertebrate Paleontology class this year: We were greatly assisted by two fantastic paleontological websites, one by Alycia Stigall at Ohio University called The Digital Atlas of Ordovician Life, and the other by Steve Holland at the University of Georgia titled The Stratigraphy and Fossils of the Upper Ordovician near Cincinnati, Ohio. Thank you to my most excellent and productive colleagues.


Brandt, D.S. 1993. Ecdysis in Flexicalymene meeki (Trilobita). Journal of Paleontology 67: 999-1005.

Brett, C.E., Thomka, J.R., Schwalbach, C.E., Aucoin, C.D. and Malgieri, T.J. 2015. Faunal epiboles in the Upper Ordovician of north-central Kentucky: Implications for high-resolution sequence and event stratigraphy and recognition of a major unconformity. Palaeoworld 24: 149-159.

Esteve, J., Hughes, N.C. and Zamora, S. 2011. Purujosa trilobite assemblage and the evolution of trilobite enrollment. Geology 39: 575-578.

Evitt, W.R. and Whittington, H.B. 1953. The exoskeleton of Flexicalymene (Trilobita). Journal of Paleontology 27: 49-55.

Foerste, A.F. 1910. Preliminary notes on Cincinnatian and Lexington fossils of Ohio, Indiana, Kentucky, and Tennessee. Denison University Science Laboratories Bulletin 16: 17-87.

Frey, R.C. 1987. The paleoecology of a Late Ordovician shale unit from southwest Ohio and southeastern Indiana. Journal of Paleontology 61: 242-267.


Wooster’s Fossil of the Week: A spherical bryozoan from the Upper Ordovician of northeastern Estonia

October 2nd, 2015

1 Esthoniopora Kukruse 585Way back in July 2007 we had our first Team Estonia doing geological field research. Andrew Milligan (’08) and I, with our friend Dr. Olev Vinn of the University of Tartu, explored the Upper Ordovician of the northeastern part of the country, perilously close to the Russian border. Most of our work was stratigraphic and related to echinoderms, but I picked up several of these beautiful spherical bryozoans. This specimen comes from the Kiviõli Member, Viivikonna Formation, Kukruse Stage, Upper Ordovician, of Kohtla-Nõmme Quarry (N 59.35665º E 27.22343º). You won’t find the quarry on a map, though, because it was soon afterwards erased by continual mining. Now it is a grassy field. Since we are studying bryozoans this week in my Invertebrate Paleontology course, I’m bringing these specimens to the blog.

2 Esthoniopora subsphaericaThis is what two specimens of this bryozoan look like before cutting. They have the size and shape of golf balls.

3 Esthoniopora subsphaericaHere are the same two specimens cut in half and polished to show the growth rings and tubular zooecia (which held the feeding zooids of the living bryozoan).

4 Esthoniopora subsphaericaIn this closer view you can see the polygonal outlines of the zooecia, now filled with calcite. In the lower right is a boring that cut through the skeleton soon after the bryozoan’s death on the Ordovician seafloor. It has a bit of sediment that filled the boring except for the very center, which apparently held the body of the borer.

This bryozoan is the trepostome Esthoniopora subsphaerica (Bassler, 1911). Bassler originally called it Hemiphragma subsphaericum, which is a nod to its abundant hemiphragms (curving partitions in the zooecial tubes). As bryozoans go, this one has a fairly simple structure with no exozone, endozone, monticules or spines. How it lived on the seafloor with such a spherical shape is a bit of a mystery. A slightly flattened patch is probably where the sphere contacted the sediment. The borings in these bryozoans were studied by Wyse Jackson and Key (2007).

5 Ray BasslerThe species author, Raymond S. Bassler (1878-1961), was an American paleontologist prominent in the study of bryozoans and other encrusting organisms. He was born in Philadelphia and became very interested in fossils from childhood. He received his bachelor’s degree from that paleontological bastion the University of Cincinnati in 1902, followed quickly by his master’s (1903) and PhD (1905) degrees from George Washington University, where he served as a professor for over forty years. He also began work at the United States National Museum in Washington in 1910, rising through the ranks to become Head Curator in 1929. His main interests were bryozoans from the Cenozoic of the Gulf and Atlantic coasts, on which he had long collaborations with the French bryozoologist Ferdinand Canu. He also worked closely with Charles Schuchert, Carl Ludwig Rominger, and Edward Oscar Ulrich. Ray Bassler died in 1961.


Bassler, R.S. 1911. The Early Paleozoic Bryozoa of the Baltic Provinces. Bulletin of the US National Museum 77: 1-382.

Koromyslova, A.V., Fedorov, P.V. and Ershova, V.B. 2009. New records of bryozoans from the Lower Ordovician of the Leningrad Region and intercolonial variability in Esthoniopora lessnikowae (Modzalevskaya). Paleontological Journal 43:39–45.

Wyse Jackson, P.N. and Key, M.M. 2007. Borings in trepostome bryozoans from the Ordovician of Estonia: two ichnogenera produced by a single maker, a case of host morphology control. Lethaia 40: 237-252.

Wooster’s Fossil of the Week: A starry bryozoan from the Upper Ordovician of southern Ohio

September 11th, 2015

Constellaria polystomella Liberty Formation 585At this time of the year I pick out one interesting specimen from the fossils my Invertebrate Paleontology class collected on their first field trip into the Upper Ordovician of southern Ohio. They did so well this week that I may be choosing a few more later! Our Fossil of the Week is the above bryozoan given the beautiful name Constellaria polystomella Nicholson, 1875. It was found by Jacob Nowell at the Caesar Creek Emergency Spillway in the Liberty Formation.
Constellaria Liberty closerConstellaria is a beautiful form, and one of the easiest bryozoans to recognize. Like all bryozooans, it was a colonial invertebrate with hundreds of filter-feeding individuals (zooids) housed in tiny tubes called zooecia. In Constellaria some of the zooecia are regularly grouped together and raised into star-shaped bumps called monticules. (The name Constellaria is clever.) This genus is a cystoporate bryozoan in the Family  Constellariidae.
JD Dana by Daniel Huntington 1858I was surprised to learn that Constellaria was named in 1846 by James Dwight Dana (1813-1895), one of the most accomplished American scientists of the 19th Century. He is best known for his Manual of Mineralogy (1848) which is still in print (greatly revised) and known as “Dana’s Mineralogy”. Dana (shown above in 1858) studied geology on scales from crystal structures to continents, with volcanoes and mountain-building in between. He had an affinity for “Zoophytes” (animals that appear to be plants), thus entangled him briefly with bryozoan systematics. Dana was born in Utica, New York, and attended Yale College, working under Benjamin Silliman, a famous chemist and mineralogist. After graduating from college he had a cool job teaching midshipmen in the US Navy, sailing through the Mediterranean in the process. For four years he served in the United States Exploring Expedition in the Pacific region. He made numerous important geological observations in Hawaii and the Pacific Northwest that he later published in books and papers. He even dabbled in theology with books like Science and the Bible: A Review of the Six Days of Creation (1856). Dana died in 1895 having received numerous accolades and awards for his research and writing.


Brown, G.D., Jr., and Daly, E.J. 1985. Trepostome bryozoa from the Dillsboro Formation (Cincinnatian Series) of southeastern Indiana. Indiana Geological Survey Special Report 33: 1-95.

Cutler, J.F. 1973. Nature of “acanthopores” and related structures in the Ordovician bryozoan Constellaria. Living and Fossil Bryozoa. Academic Press, London, 257-260.

Dana J.D. 1846. Structure and classification of zoophytes. U.S. Exploring Expedition 1838-1842, 7: 1-740.

Wooster Paleontologists return to Caesar Creek Lake

September 6th, 2015

1 Shoreside collecting 090615Ohio is a wonderful place for paleontologists. One of the reasons is the thick, productive set of Upper Ordovician rocks that are exposed in the southwest of the state in and around Cincinnati. It is an easy drive south from Wooster into some of the most fossiliferous sediments in the world. Today Wooster’s Invertebrate Paleontology class went to Caesar Creek Lake in Warren County with its shoreline and emergency spillway exposing richly productive limestones and shales of the Waynesville, Liberty and Whitewater Formations (all of which equal the Bull Fork Formation). I’ve been there many times with many classes. The weather today was hot and dry — a contrast with last year’s torrential rains and mud.

2 Caesar Creek Lake Visitor Center MS 090615After a three-hour drive, we stopped first at the U.S. Army Corps of Engineers visitor center for Caesar Creek Lake. The Corps built the dam and spillway for the lake, and continues to maintain them both. This center has a set of museum displays and is where we obtain our fossil collecting permit. (This image was taken by Mara Sheban, a sophomore at The College of Wooster who will be contributing photographs to this blog and other geology projects.)

3 Initial briefing MS 090615Our initial briefing, with a review of the local geology and most common fossils. Since this is the start of the Invertebrate Paleontology course, most students have only begun to explore the topic. The fossils they collect on this trip will be the basis of a semester-long project of systematics and paleoecological analysis. (Photo by Mara Sheban.)

4 Marching to the outcrop 090615Our march to the far north of the exposures at the Caesar Creek Lake emergency spillway. We actually drop down from the spillway to the lake’s south shore. (See photo at the top of this post.) In the distant parking lot you can just make out a white pick-up truck in which the seasonal ranger is keeping an eye on us.

5 Jacob Nowell collecting MS 090615Jacob Nowell collecting small fossils washed from the shale along the lakeshore. (Photo by Mara Sheban.)

6 Jacob Pries trilobite MS 090615Jacob Pries found a nice enrolled Flexicalymene trilobite. Caesar Creek has a reputation as being a good place to find trilobites. We love them, but are also interested in the rest of the fossil fauna. (Mara Sheban image.)

7 Brachiopod hash 090615Some of the limestone units are a nearly solid hash of cemented brachiopod shells.

8 Hardground full view 0090616My favorite slabs at Caesar Creek Lake are the abundant carbonate hardgrounds like the above. A hardground is a cemented seafloor, usually with borings and encrusting organisms. This one began as a burrowed soft carbonate sediment. The burrows were filled with fine mud that cemented early on the seafloor. The surrounding softer matrix washed away, leaving a hardground with the burrows now in positive relief. Brachiopods and corals then lived in the nooks and crannies of this hard rock on the bottom of the sea.

9 Hardground closer 090615Here is the burrow system in a closer view. In the upper right is a beautiful encrusted rugose coral, shown in detail below.

10 Bryo encrusted rugosan 090615That coral is almost completely covered by a trepostome bryozoan. I would have loved to collect this specimen for further study, but the slab is too large and no tools are allowed on this outcrop.

In the coming weeks we will identify the fossils we gathered, apply to them several paleontological techniques such as cleaning, cutting, polishing and photography, and then put together a grand paleoecological analysis. We will be greatly assisted by two fantastic websites, one by Alycia Stigall at Ohio University called The Digital Atlas of Ordovician Life, and the other by Steve Holland at the University of Georgia titled The Stratigraphy and Fossils of the Upper Ordovician near Cincinnati, Ohio.

Wooster’s Fossil of the Week: Small and common orthid brachiopods from the Upper Ordovician of Ohio

August 7th, 2015

Cincinnetina meeki (Miller, 1875) slab 1 585
One of the many benefits of posting a “Fossil of the Week” is that I learn a lot while researching the highlighted specimens. I not only learn new things, I learn that some things I thought I knew must be, shall we say, updated. The above slab contains dozens of brachiopods (and a few crinoid ossicles and bryozoans). I have long called the common brachiopod here Onniella meeki. Now I learn from my colleagues Alycia Stigall and Steve Holland at their great Cincinnatian websites that since 2012 I should be referring to this species as Cincinnetina meeki (Miller, 1875). Jisuo Jin sorted out its taxonomy in a Palaeontology article three years ago:

Phylum: Brachiopoda
Class: Rhynchonellata
Order: Orthida
Family: Dalmanellidae
Genus: Cincinnetina
Species: Cincinnetina meeki (Miller, 1875)
Cincinnetina meeki (Miller, 1875) slab 2 585This slab, which resides in our Geology 200 teaching collection, was found at the famous Caesar Creek locality in southern Ohio. It is from the Waynesville/Bull Fork Formation and Richmondian (Late Ordovician) in age.
Cincinnetina meeki (Miller, 1875) slab 3 585You may see some bryozoans in this closer view. This bed is a typical storm deposit in the Cincinnatian Group. The shells were tossed about, most landing in current-stable conditions, and finer sediments were mostly washed away, leaving this skeletal lag.


Jin, J. 2012. Cincinnetina, a new Late Ordovician dalmanellid brachiopod from the Cincinnati type area, USA: implications for the evolution and palaeogeography of the epicontinental fauna of Laurentia. Palaeontology 55: 205–228.

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