Wooster’s Fossil of the Week: A common trilobite from the Upper Ordovician of Ohio

December 11th, 2015

Flexicalymene meeki cephalon view 585This beautiful specimen was collected by Wooster student Eve Caudill on this year’s College of Wooster Invertebrate Paleontology field trip to Caesar Creek Lake, Ohio. It is the iconic trilobite Flexicalymene meeki (Foerste, 1910) from a soft, “buttery” shale in the Waynesville Formation (Upper Ordovician). This is one of the most common trilobite species in the world, and it has been photographed thousands of times, so I posed it at an unconventional, rakish angle. We are looking here at the cephalon (head) of the animal. I like the way the remnants of the enclosing sedimentary matrix cling to the low places, highlighting the bumps and ridges. The center of the cephalon shows the distinctive glabella with its side lobes. The stomach of the trilobite was housed underneath it. The two eyes are visible on either side of the glabella, the one on the right split by the slightly-open facial suture used for dividing its exoskeleton during molting (ecdysis).

Flexicalymene meeki pygidium view 585This is a view of the pygidium (tail end) of the same Flexicalymene meeki specimen. It is tucked under the leading edge of the cephalon in the classic enrollment position. Trilobites likely enrolled for several reasons, but the primary one was almost certainly to affect a pill-bug-like defense against predators.

Flexicalymene meeki side view 585This is a side view of the enrolled trilobite. The articulated segments between the cephalon and pygidium constitute the thorax.foerste-1936We met the author of Flexicalymene meeki four years ago in this blog, so let’s visit him again. August F. Foerste (1862-1936) was one of the pioneers of Cincinnatian paleontology and stratigraphy. He grew up and worked in the Dayton, Ohio, area. Foerste went to Denison University where he was a very successful undergraduate, publishing several geological papers. He returned to Dayton after graduation with a PhD from Harvard, teaching high school for 38 years. When he retired he was offered a teaching position at the University of Chicago, but instead went to work at the Smithsonian Institution until the end of his life.

A final note from the Invertebrate Paleontology class this year: We were greatly assisted by two fantastic paleontological websites, one by Alycia Stigall at Ohio University called The Digital Atlas of Ordovician Life, and the other by Steve Holland at the University of Georgia titled The Stratigraphy and Fossils of the Upper Ordovician near Cincinnati, Ohio. Thank you to my most excellent and productive colleagues.


Brandt, D.S. 1993. Ecdysis in Flexicalymene meeki (Trilobita). Journal of Paleontology 67: 999-1005.

Brett, C.E., Thomka, J.R., Schwalbach, C.E., Aucoin, C.D. and Malgieri, T.J. 2015. Faunal epiboles in the Upper Ordovician of north-central Kentucky: Implications for high-resolution sequence and event stratigraphy and recognition of a major unconformity. Palaeoworld 24: 149-159.

Esteve, J., Hughes, N.C. and Zamora, S. 2011. Purujosa trilobite assemblage and the evolution of trilobite enrollment. Geology 39: 575-578.

Evitt, W.R. and Whittington, H.B. 1953. The exoskeleton of Flexicalymene (Trilobita). Journal of Paleontology 27: 49-55.

Foerste, A.F. 1910. Preliminary notes on Cincinnatian and Lexington fossils of Ohio, Indiana, Kentucky, and Tennessee. Denison University Science Laboratories Bulletin 16: 17-87.

Frey, R.C. 1987. The paleoecology of a Late Ordovician shale unit from southwest Ohio and southeastern Indiana. Journal of Paleontology 61: 242-267.


Wooster’s Fossil of the Week: A fragmentary rostroconch from the Middle Devonian of Ohio

November 27th, 2015

1 Hippocardia 1Not all of our featured fossils are particularly beautiful, or even entire, but they are interesting in some way. Above is the broken cross-section of a rostroconch mollusk known as Hippocardia Brown, 1843. It was found somewhere in Ohio by the late Keith Maneese and kindly donated to the department by his widow Cameron Maneese. From its preservation and the kind of rock making up the matrix inside, we can tell that it almost certainly came from the Columbus Limestone (Middle Devonian, Eifelian).

In the top image it is apparent that this fossil has bilateral symmetry, a heart-shaped cross-section, and a ribbed calcitic shell. This is the dorsal view.
2 Hippocardia 2Flipping the specimen upside-down, we now have a view of the ventral portion. Again we see the ribs and bilateral symmetry.
3 Hippocardia side viewThis side view shows that the ribs extend from the dorsal to the ventral sides and are angled to the axis of the shell. That’s about all we can tell. (And this is the best specimen of a rostroconch we have! Thank you again, Cameron.)
4 CzechVirtualRostroThis diagram of a complete rostroconch (from the Czech Virtual Museum). This is a side view of a species that does not have the dorsal-ventral ribbing. The shell is superficially like that of a bivalve (clam), but the valves are fused together and their is a distinctive tube (rostrum) extending to the posterior. Much study and debate about the rostroconchs has at least confirmed that these are a class of mollusks separate from the bivalves. They lived semi-infaunally with the rostrum extending into the seawater to channel a flow of water into the body chamber for filter-feeding, much like infaunal bivalves today that have siphons. Rostroconchs and cephalopods appear to be sister groups, and some rostroconchs may have evolved into the scaphopods. Plenty of arguments to go around, though, on the evolution and diversification of mollusks
5 Thomas 1843 p 976 Thomas pl 8 fig 10 1843Captain Thomas Brown (1785-1862) named the genus Hippocardia in 1843. He was a Scottish naturalist who studied many topics, including mollusks. Above are the sections of his book The Elements of Fossil Conchology that describe and illustrate Hippocardia (considering it a bivalve). Captain Brown was born in Perth and went to school in Edinburgh. He joined the militia at 20, becoming a captain at 26. When he was transferred to Manchester, England, Brown acquired an interest in nature. He bought a flax mill after leaving the military, but it burned down while still uninsured. He thus turned to nature writing for support. He was became a Fellow of the Royal Society of Edinburgh in 1818, and in 1840 he was appointed curator of the Manchester Museum. He retained this position for the rest of his life. He was later a Fellow of the Linnean Society and a member, in classic 19th Century fashion, of several other groups, including the Wernerian, Kirwanian and Phrenological Societies. (I love the addition of phrenology to his interests!) The marine gastropod Zebina browniana d’Orbigny, 1842, was named after him. An interesting character, this Captain Brown, but I’ve been unable to find a single portrait of him.


Brown, T. 1843. The elements of fossil conchology according to the arrangement of Lamarck; with the newly established genera of other authors. Houlston & Stoneman, London; 133 pages.

Hoare, R.D. 1989. Taxonomy and paleoecology of Devonian rostroconch mollusks from Ohio. Journal of Paleontology 63: 838-846.

Pojeta, J., Jr., Runnegar, B. 1976. The paleontology of rostroconch mollusks and the early history of the phylum Mollusca. United States Geological Survey Professional Paper 968: 1-88.

Pojeta, J., Jr., Runnegar, B., Morris, N.J. and Newell, N.D. 1972. Rostroconchia: a new class of bivalved mollusks. Science 177: 264-267.

Runnegar, B., Goodhart, C.B. and Yochelson, E.L. 1978. Origin and evolution of the Class Rostroconchia [and discussion]. Philosophical Transactions of the Royal Society B: Biological Sciences 284(1001): 319-333.

Wagner, P.J. 1997. Patterns of morphologic diversification among the Rostroconchia. Paleobiology 23: 115-150.

Wooster’s Pseudofossils of the Week: Cone-in-cone structures from southern Ohio

October 30th, 2015

1 ShatterCones 585

Author’s note: James Chesire convinced me through the comments and later correspondence that what we actually have here are cone-in-cone structures, not shatter cones. I’ve thus changed the title but have left the post below in its original form. They are still pseudofossils. I’ll link here later for a full update. Thanks, James!

This complex rock was collected decades ago in Adams County, Ohio, by the late Professor Frank L. Koucky of The College of Wooster. He was at the time studying a strange geological feature in that part of the state known then as the Serpent Mound Cryptoexplosion Structure. He thought that the ring-like disturbance in the bedrock nearly 10 km wide was a place where “mantle gases” explosively erupted from below. The rock shown was going to be a key to deciphering the energy of these cataclysmic events. It is a set of shatter cones formed when enormous, high velocity pressures were applied to a micritic (fine-grained) limestone. Professor Koucky knew what these features represented, but they are still collected in that region and elsewhere as “fossils” by some because of their resemblance to corals. They are thus fine examples of pseudofossils, or inorganic features resembling fossils.

These shatter cones ended up showing conclusively that the event that caused the “bedrock disturbance” in southern Ohio was actually an ancient meteor impact, and the site is now known as the Serpent Mound Crater. This ancient crater (it may be as much as 320 million years old) has a central uplift surrounded by a ring graben (circular down-dropped rocks). It took a lot of clever geology to sort this out because known of it is now visible on the surface.
2 Shatter cones closerThe Serpent Mound shatter cones have a multiple long fractures running parallel to the cones, resembling hair or “horsetails”. The cones have horizontal step-like fractures on their broken surfaces. You can simulate this kind of structure by firing a BB or small rock at thick glass, which produces a conical fracture and perpendicular steps. To do this in a limestone requires between 20 and 200 kbar of pressure, which can only be achieved by a large meteorite impact or a nuclear explosion underground. More likely it was the former!
3 Shatter cones plan viewHere is what these shatter cones look like in plan view. The hole in the upper left is the tip of a cone that is not preserved.

So, shatter cones, despite their fine and repeatable details, are inorganic and not fossils of any kind. They represent enormous shock waves that left their marks as they passed through this limestone many millions of years ago.


Carlton, R.W., Koeberl, C., Baranoski, M.T. and Schumacher, G.A. 1998. Discovery of microscopic evidence for shock metamorphism at the Serpent Mound structure, south-central Ohio: confirmation of an origin by impact. Earth and Planetary Science Letters 162: 177-185.

Kenkmann, T., Poelchau, M.H., Trullenque, G., Hoerth, T., Schäfer, F., Thoma, K. and Deutsch, A. 2012. Shatter cones formed in a MEMIN impact cratering experiment. Meteoritics and Planetary Science Supplement 75: 5092.

Milton, D.J. 1977. Shatter cones – an outstanding problem in shock mechanics. In: Impact and Explosion Cratering: Planetary and Terrestrial Implications 1: 703-714.

Sagy, A., Fineberg, J. and Reches, Z. 2004. Shatter cones: Branched, rapid fractures formed by shock impact. Journal of Geophysical Research 109: B10209.

Wooster’s Fossil of the Week: A starry bryozoan from the Upper Ordovician of southern Ohio

September 11th, 2015

Constellaria polystomella Liberty Formation 585At this time of the year I pick out one interesting specimen from the fossils my Invertebrate Paleontology class collected on their first field trip into the Upper Ordovician of southern Ohio. They did so well this week that I may be choosing a few more later! Our Fossil of the Week is the above bryozoan given the beautiful name Constellaria polystomella Nicholson, 1875. It was found by Jacob Nowell at the Caesar Creek Emergency Spillway in the Liberty Formation.
Constellaria Liberty closerConstellaria is a beautiful form, and one of the easiest bryozoans to recognize. Like all bryozooans, it was a colonial invertebrate with hundreds of filter-feeding individuals (zooids) housed in tiny tubes called zooecia. In Constellaria some of the zooecia are regularly grouped together and raised into star-shaped bumps called monticules. (The name Constellaria is clever.) This genus is a cystoporate bryozoan in the Family  Constellariidae.
JD Dana by Daniel Huntington 1858I was surprised to learn that Constellaria was named in 1846 by James Dwight Dana (1813-1895), one of the most accomplished American scientists of the 19th Century. He is best known for his Manual of Mineralogy (1848) which is still in print (greatly revised) and known as “Dana’s Mineralogy”. Dana (shown above in 1858) studied geology on scales from crystal structures to continents, with volcanoes and mountain-building in between. He had an affinity for “Zoophytes” (animals that appear to be plants), thus entangled him briefly with bryozoan systematics. Dana was born in Utica, New York, and attended Yale College, working under Benjamin Silliman, a famous chemist and mineralogist. After graduating from college he had a cool job teaching midshipmen in the US Navy, sailing through the Mediterranean in the process. For four years he served in the United States Exploring Expedition in the Pacific region. He made numerous important geological observations in Hawaii and the Pacific Northwest that he later published in books and papers. He even dabbled in theology with books like Science and the Bible: A Review of the Six Days of Creation (1856). Dana died in 1895 having received numerous accolades and awards for his research and writing.


Brown, G.D., Jr., and Daly, E.J. 1985. Trepostome bryozoa from the Dillsboro Formation (Cincinnatian Series) of southeastern Indiana. Indiana Geological Survey Special Report 33: 1-95.

Cutler, J.F. 1973. Nature of “acanthopores” and related structures in the Ordovician bryozoan Constellaria. Living and Fossil Bryozoa. Academic Press, London, 257-260.

Dana J.D. 1846. Structure and classification of zoophytes. U.S. Exploring Expedition 1838-1842, 7: 1-740.

Wooster Paleontologists return to Caesar Creek Lake

September 6th, 2015

1 Shoreside collecting 090615Ohio is a wonderful place for paleontologists. One of the reasons is the thick, productive set of Upper Ordovician rocks that are exposed in the southwest of the state in and around Cincinnati. It is an easy drive south from Wooster into some of the most fossiliferous sediments in the world. Today Wooster’s Invertebrate Paleontology class went to Caesar Creek Lake in Warren County with its shoreline and emergency spillway exposing richly productive limestones and shales of the Waynesville, Liberty and Whitewater Formations (all of which equal the Bull Fork Formation). I’ve been there many times with many classes. The weather today was hot and dry — a contrast with last year’s torrential rains and mud.

2 Caesar Creek Lake Visitor Center MS 090615After a three-hour drive, we stopped first at the U.S. Army Corps of Engineers visitor center for Caesar Creek Lake. The Corps built the dam and spillway for the lake, and continues to maintain them both. This center has a set of museum displays and is where we obtain our fossil collecting permit. (This image was taken by Mara Sheban, a sophomore at The College of Wooster who will be contributing photographs to this blog and other geology projects.)

3 Initial briefing MS 090615Our initial briefing, with a review of the local geology and most common fossils. Since this is the start of the Invertebrate Paleontology course, most students have only begun to explore the topic. The fossils they collect on this trip will be the basis of a semester-long project of systematics and paleoecological analysis. (Photo by Mara Sheban.)

4 Marching to the outcrop 090615Our march to the far north of the exposures at the Caesar Creek Lake emergency spillway. We actually drop down from the spillway to the lake’s south shore. (See photo at the top of this post.) In the distant parking lot you can just make out a white pick-up truck in which the seasonal ranger is keeping an eye on us.

5 Jacob Nowell collecting MS 090615Jacob Nowell collecting small fossils washed from the shale along the lakeshore. (Photo by Mara Sheban.)

6 Jacob Pries trilobite MS 090615Jacob Pries found a nice enrolled Flexicalymene trilobite. Caesar Creek has a reputation as being a good place to find trilobites. We love them, but are also interested in the rest of the fossil fauna. (Mara Sheban image.)

7 Brachiopod hash 090615Some of the limestone units are a nearly solid hash of cemented brachiopod shells.

8 Hardground full view 0090616My favorite slabs at Caesar Creek Lake are the abundant carbonate hardgrounds like the above. A hardground is a cemented seafloor, usually with borings and encrusting organisms. This one began as a burrowed soft carbonate sediment. The burrows were filled with fine mud that cemented early on the seafloor. The surrounding softer matrix washed away, leaving a hardground with the burrows now in positive relief. Brachiopods and corals then lived in the nooks and crannies of this hard rock on the bottom of the sea.

9 Hardground closer 090615Here is the burrow system in a closer view. In the upper right is a beautiful encrusted rugose coral, shown in detail below.

10 Bryo encrusted rugosan 090615That coral is almost completely covered by a trepostome bryozoan. I would have loved to collect this specimen for further study, but the slab is too large and no tools are allowed on this outcrop.

In the coming weeks we will identify the fossils we gathered, apply to them several paleontological techniques such as cleaning, cutting, polishing and photography, and then put together a grand paleoecological analysis. We will be greatly assisted by two fantastic websites, one by Alycia Stigall at Ohio University called The Digital Atlas of Ordovician Life, and the other by Steve Holland at the University of Georgia titled The Stratigraphy and Fossils of the Upper Ordovician near Cincinnati, Ohio.

A Wooster Geologist goes to a Bigfoot meeting

August 29th, 2015

1 Bigfoot head reconstruction 082915ORRVILLE, OHIO — The First-Year Seminar course I teach at Wooster is called “Nonsense! (And Why it’s So Popular)“. It is ostensibly about exploring irrational ideas in human society, such as astrology, conspiracy theories, pseudoscience, quack medicine, the “paranormal” and the like, but more fundamentally concerned with critical thinking and writing. It is about skepticism and learning how to test ideas and express the results. It is great fun because there is of course an endless carousel of nonsense to choose from every semester. We’re careful not to ridicule people, but we assess myths and misconceptions ruthlessly. Understanding why people believe weird things (from the title of one of our textbooks) turns out to be just as interesting as the ideas themselves, and it reveals the many filters and barriers between us and “reality” or “truth”. There are fuzzy boundaries around every topic, and our empathy for people who have poorly-supported world views grows throughout the course. Still, we can call some concepts nonsense even if the people professing them are sympathetic. The existence of Bigfoot is unsupported by physical evidence. The claim that Bigfoot has been thriving in Ohio for thousands of years is utter nonsense.

So why do people believe that an eight-foot tall hominid has been hiding all this time in the woods and swamps of crowded Ohio? To gain some insight, today my wife Gloria and I went to a Bigfoot presentation held at the Orrville Public Library by the founders of American Primate Exploration (APE from now on!). It was a fascinating two hours. You can see what the primary program was like by watching this video made from an earlier but nearly identical event.

2 Bigfoot presentation 082915The main presenter was Dan Baker, pictured above, the founder of APE. He first described the history and organization of his organization, emphasizing the number of “research teams” they have across North America and even into Australia. All the teams are staffed by what he termed “self-described experts on Bigfoot”, then noting that no one is a real expert on Bigfoot (thus leveling the field with scientists, I suppose). Mr. Baker moved immediately into anatomy, showing how Bigfoot has a flexible foot structure that includes a “midtarsal break”, unlike most humans. This means that Bigfoot footprints show a distinctive pressure ridge behind the ball of the foot, separating them from the typical human footprint. He had to admit, though, that one in 13,000 humans has such a mid-tarsal break. Turns out it is actually one in 13, so he was three orders of magnitude off. [Update: Check out the “midtarsal break” on these fake Bigfoot tracks.]

3 Bigfoot footprint casts 082915Above is a selection of the footprint casts displayed at the meeting. As you may can see, sorting out evidence for a midtarsal break from a regular human-like arch is dodgy business. Plus, some of those Bigfoot impressions are flat throughout, a feature attributed to “casting errors” by one of the presenters.

I was amazed to see that the famous 1967 Patterson-Gimlin film of a “Bigfoot” now named “Patty” was a central piece of “evidence” for the existence of Bigfoot. Mr. Baker breezily dismissed all accusations of fraud on the part of Patterson and Gimlin, which include a confession from the man filmed in the ape suit. This ancient clip, it turns out, is now holy script in the Bigfoot movement, so no skeptical analysis will make a dent in this belief system. Mr. Baker even brought a footprint cast that he proudly showed was signed by Bob Gimlin himself. A sacred relic.

4 Bigfoot handprint 082915I was introduced to evidence new to me: Bigfoot handprints. Here is one that was apparently made in 1995 by Paul Freeman in the Blue Mountains, Washington. Note the opposable thumb. Pretty impressive how that animal pushed his whole hand down into the mud. (The fingers are shortened here because, one of the APE crew said, “he didn’t clean out the holes first”.) Mr. Baker said that this cast all all the work of Paul Freeman was “legit”, despite claims that the man was a fraud. You can follow up on the Paul Freeman story here: “A few of Freeman’s “Bigfoot-related” discoveries were found to be faked, including manmade hair samples, and a few of his finds remain “unknowns.” (Although softened, this criticism is heresy in Bigfoot circles.) Freeman’s most famous film of a Bigfoot was shown approvingly at our event. Freeman had an unusual ability to find Bigfoot, proponents say. Unusual indeed.

After these items, Mr. Baker and the following speaker (Raymond V. Gardner II, APE Field Researcher) spent the remaining time describing their own encounters with Bigfoot in Ohio. Carroll County (about an hour and a half east of Wooster) is a hotspot, as is the area around Spencer in Medina County (just a half hour up the road). In fact, Ohio itself is second only to the Pacific Northwest in Bigfoot reports. The stories were what you would expect: fleeting glimpses before a camera could be deployed, howls and “wood-banging” in the night, “trampled grass” after a night of “activity”. One long segment was an audio recording of an APE team describing some squatting shape in the midnight woods. There was a brief mention of giving some vocal Bigfoot recordings to “a Native American” who could apparently translate them into some sort of pidgin English. (I’ve never heard that before.) I also learned that Bigfoot may be able to “see in the infrared” and thus avoid the ubiquitous trail cameras in the Ohio woods.

The questions from the audience were interesting. Most there were true believers from their tone, but some skeptics lurked. One of the best questions asked why we don’t see evidence of Bigfoot in the Pacific Northwest right now as unprecedented wildfires tear through the forests. Shouldn’t Bigfoot be flushed out into the open, or at least a few smoking bodies be found? No, was the answer. Bigfoot is very smart, very crafty, very quick, and very good at hiding. The lack of evidence is not evidence that they don’t exist. I also learned from an audience member that Bigfoot sometimes speaks in “backwards Indian” and “the Douglas Dialect“.

My conclusion is that we attended a service with true Bigfoot believers. The lack of evidence for the creature is quickly explained away to preserve the tenets of the faith. Bigfoot proponents have invested their identities in its existence, no matter how implausible. No amount of scientific skepticism can overcome a belief unencumbered by a need for physical evidence or even biological possibility. Bigfoot believers have a strong community reinforced daily with testimonies and acts of resistance against skeptics. As with any community, there are social roles to fill, from the leadership to “field researchers”. There is even an Ohio Bigfoot Hall of Fame for the ambitious. If your subject is defined by perpetual ambiguity, your arguments for it can be impervious to logical analysis. This was an excellent field experience for me to bring back to my First-Year Seminar class next week.

Thank you to the good and earnest officers of APE for the presentation (free to the public), and to the Orrville Public Library for hosting the event.

Here’s a cool book to explore the origins and details of Bigfoot mythology —

618c3NRVZML._SX329_BO1,204,203,200_UPDATE: Excellent blog post from Scientific American on the implausibility of Bigfoot’s existence.

Wooster’s Fossil of the Week: Small and common orthid brachiopods from the Upper Ordovician of Ohio

August 7th, 2015

Cincinnetina meeki (Miller, 1875) slab 1 585
One of the many benefits of posting a “Fossil of the Week” is that I learn a lot while researching the highlighted specimens. I not only learn new things, I learn that some things I thought I knew must be, shall we say, updated. The above slab contains dozens of brachiopods (and a few crinoid ossicles and bryozoans). I have long called the common brachiopod here Onniella meeki. Now I learn from my colleagues Alycia Stigall and Steve Holland at their great Cincinnatian websites that since 2012 I should be referring to this species as Cincinnetina meeki (Miller, 1875). Jisuo Jin sorted out its taxonomy in a Palaeontology article three years ago:

Phylum: Brachiopoda
Class: Rhynchonellata
Order: Orthida
Family: Dalmanellidae
Genus: Cincinnetina
Species: Cincinnetina meeki (Miller, 1875)
Cincinnetina meeki (Miller, 1875) slab 2 585This slab, which resides in our Geology 200 teaching collection, was found at the famous Caesar Creek locality in southern Ohio. It is from the Waynesville/Bull Fork Formation and Richmondian (Late Ordovician) in age.
Cincinnetina meeki (Miller, 1875) slab 3 585You may see some bryozoans in this closer view. This bed is a typical storm deposit in the Cincinnatian Group. The shells were tossed about, most landing in current-stable conditions, and finer sediments were mostly washed away, leaving this skeletal lag.


Jin, J. 2012. Cincinnetina, a new Late Ordovician dalmanellid brachiopod from the Cincinnati type area, USA: implications for the evolution and palaeogeography of the epicontinental fauna of Laurentia. Palaeontology 55: 205–228.

Wooster’s Fossil of the Week: A coiled nautiloid from the Middle Devonian of Ohio

July 17th, 2015

Goldringia cyclops Columbus Ls Devonian 585The above fossil is a nautiloid cut in cross-section, showing the large body chamber at the bottom and behind it to the left and above the phragmocone, or chambered portion of the conch (shell). It is a species of Goldringia Flower, 1945, found in the Columbus Limestone (Middle Devonian, Eifelian) exposed in the Owen Stone Quarry near Delaware, Ohio. It is a nice specimen for both what it shows us about a kind of nautiloid coiling and for clues to its preservation.

This specimen was originally labelled Gyroceras cyclops Hall, 1861. In 1945, Rousseau Flower designated this taxon the type species of Goldringia. I can’t tell if we really have G. cyclops here or some other species, so I’m leaving it at the genus level. The old name lingers, though, in the term for this kind of open coiling: gyroceraconic. It is one of the earliest examples of the nautiloids having the phragmocone positioned above the body chamber, presumably for stable buoyancy.
Pentamerid embedded 071315I like the clues to the early history of this conch after death. The chambers are entirely filled with sediment, a fossiliferous micrite. You can see places where the original shell was broken and larger bits infiltrated, like the whole brachiopod shown above. This brachiopod appears from its cross-section to be a pentamerid. Also visible are strophomenid brachiopods and gastropods.
Winifred GoldringRousseau Hayner Flower (1913–1988) described Goldringia in 1945. He doesn’t directly say who he named it after, but he thanks “Dr. Winifred Goldring of the New York State Museum” in the acknowledgments. We can tell Flower’s story later (and it’s a good one), but this gives us a chance to introduce Winifred Goldring (1888-1971). She was the first paleontologist to describe the famous Gilboa fossil flora (Devonian) in upstate New York, and she was the first woman State Paleontologist of New York (or anywhere, for that matter). (Now there is Lisa Amati in this prestigious position. Congratulations, Lisa!) Goldring grew up near Albany, New York, one of nine children in a very botanical family. She graduated from Wellesley College in 1909 with a bachelor’s degree in geology (very unusual for a woman at the time). She stayed at Wellesley to earn a master’s degree (1912). She also taught geology courses at Wellesley. In 1913 she studied geology at Columbia University with the famous Amadeus Grabau. In 1914, Goldring joined the scientific staff at the New York State Museum as a “scientific expert”. She worked her way up through the many ranks there to become State Paleontologist in 1939. She is best known as a paleontologist for her work with the fascinating Gilboa fossil forest, bringing her early upbringing by botanists to full circle. Along the way she was the first woman president of the Paleontological Society (in 1949) and vice-president of the Geological Society of America (in 1950). A hero of paleontology.


Flower, R.H. 1945. Classification of Devonian nautiloids. American Midland Naturalist 33: 675–724.

Goldring, W. 1927. The oldest known petrified forest. Scientific Monthly 24: 514–529.

Koninck, L.G.D. 1880. Faune du Calcaire Carbonifere de la Belgique, deuxieme partie, Genres Gyroceras, Cyrtoceras, Gomphoceras, Orthoceras, Subclymenia et Goniatites. Annales du Musee Royal d‘Histoire Naturelle, Belgique 5: 1–333.

Wooster’s Fossils of the Week: A Silurian encrinite from southwestern Ohio

May 22nd, 2015

BrassfieldEncrinite585_041915The above rock was collected on our Sedimentology & Stratigraphy class field trip last month. It is an average piece of weathered Brassfield Formation (Early Silurian, Llandovery) from Oakes Quarry Park near Fairborn, Ohio (N 39.81472°, W 83.99471°). It is made almost entirely of crinoid fragments, and has a pleasant pinkish hue, most of which comes from the crinoid bits themselves. If you look closely you can see crinoid thecal plate fragments as well columnals and pluricolumnals.

This kind of limestone in which echinoderm ossicles make up the bulk of the grains is known as an encrinite. I first learned about encrinites from my colleague Bill Ausich of The Ohio State University, who has written the best assessments of encrinites on a regional scale. Encrinites are well-washed biosparite grainstones typically deposited between fair weather and storm wave bases on shallow shelves in low latitudes. They are surprisingly common from the Ordovician into the Jurassic, but then the disappear from the rock record as crinoids declined in abundance in shallow environments.

We’ve seen encrinites before in this blog from the Silurian of Estonia, the Triassic of Poland, and the Jurassic of Utah.


Ausich, W.I. 1986. Early Silurian inadunate crinoids (Brassfield Formation, Ohio). Journal of Paleontology 60: 719-735.

Ausich, W.I. 1997. Regional encrinites: a vanished lithofacies. In: Paleontological events: stratigraphic, ecologic and evolutionary implications, p. 509-519. Columbia University Press, New York.

Ausich, W.I. and Deline, B. 2012. Macroevolutionary transition in crinoids following the Late Ordovician extinction event (Ordovician to Early Silurian). Palaeogeography, Palaeoclimatology, Palaeoecology 361: 38-48.

Hunter, A.W. and Zonneveld, J.P. 2008. Palaeoecology of Jurassic encrinites: reconstructing crinoid communities from the Western Interior Seaway of North America. Palaeogeography, Palaeoclimatology, Palaeoecology 263: 58-70.

Tang, C.M., Bottjer, D.J. and Simms, M.J. 2000. Stalked crinoids from a Jurassic tidal deposit in western North America. Lethaia 33: 46-54.

Wooster’s Fossil of the Week: A ptilodictyine bryozoan from the Silurian of Ohio

May 15th, 2015

Phaenopora superba Brassfield 585The fossil above was found by Luke Kosowatz (’17) on our Sedimentology & Stratigraphy class field trip last month. We were measuring and sampling the Brassfield Formation (Early Silurian, Llandovery) near Fairborn, Ohio, and Luke pulled this beauty out of the rubble. This limestone is full of echinoderms and corals, so this lonely bryozoan was immediately a star.
Peela 050815This is the specimen that we sectioned and made an acetate peel from last month. The interior view, shown above, was critical to its identification. This peel was made perpendicular to the surface. It shows that the bryozoan is bifoliate, meaning it has two sides with zooids (the filter-feeding bryozoan polypides) and stood upright on the seafloor like a fan or leaf. Both sides had the characteristic bumps called monticules.
Phaenopora closerThe next critical view is this close-up of a slightly weathered surface of the bryozoan. It shows a regular arrangement of the larger zooecia (autozooecia) with two smaller zoooecia (metazooecia) between each pair. These clues enabled my friend Andrej Ernst, a paleontologist and bryozoan expert in the Department of Geosciences at the University of Hamburg, to identify this bryozoan as the ptilodictyine Phaenopora superba (Billings, 1866).
CNSPhoto-GEOLOGISTElkanah Billings (1820-1876) originally described this bryozoan species in 1866. He was Canada’s first government paleontologist, and he very much looked the part. Billings was born on a farm near Ottawa. He went to law school and became a lawyer in 1845, but he gave up dusty books for the life of a field paleontologist. In 1856 Billings joined the Geological Survey of Canada. He named over a thousand new species in his career. The Billings Medal is given annually by the Geological Association of Canada to the most outstanding of its paleontologists.


Billings, E. 1866. Catalogues of the Silurian fossils of the island of Anticosti: with descriptions of some new genera and species. Dawson brothers.

Ross, J.P. 1960. Larger cryptostome Bryozoa of the Ordovician and Silurian, Anticosti Island, Canada: Part I. Journal of Paleontology 34: 1057-1076.

Ross, J.P. 1961. Larger cryptostome Bryozoa of the Ordovician and Silurian, Anticosti Island, Canada: Part II. Journal of Paleontology 35: 331-344.

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