Wooster’s Fossil of the Week: a twisted little crinoid (Lower Silurian of Estonia)

September 9th, 2012

This week’s fossil is a tiny little crinoid with an odd shape. Calceocrinus balticensis (shown above with the scale bar as one millimeter) is a new species from the Lower Silurian (Llandovery) of Hiiumaa, western Estonia. It is part of a series of new crinoid taxa described in the most recent issue of Acta Palaeontologica Polonica by Ausich et al. (2012). All that geological work in Estonia by Ohio State and Wooster geologists is resulting in several paleontological publications, all with the collaboration of our friend Olev Vinn at the University of Tartu, Estonia.

The western Estonian island of Hiiumaa where our little crinoid was found. (Image courtesy of Google Maps.)

Calceocrinus balticensis Ausich, Wilson and Vinn, 2012 (to give its full and glorious name) is unusual because its crown (the filter-feeding “head” of the crinoid) is recumbent on the column (the “stem”). In the images above you can see the column as a series of disks on their sides at the bottom of the view. The crown is the set of larger plates attached to the top of the column, from which there are several arms extending to the right. This new species is the first of its genus from the paleocontinent Baltica. It had sister species in North America on what became Anticosti Island in eastern Canada (see Ausich and Copper, 2010).

Calceocrinids (Order Calceocrinida Ausich, 1998) lived very close to the seafloor. The column of an individual, which in other crinoids holds the crown far off the substrate, lay horizontally along the bottom. The crown was hinged at its base so that it could be elevated perpendicular to the stem with the arms spread wide to filter organic material from the water. During non-feeding times the crown would lie inconspicuous on the bottom. This crinoid literally had a very low profile compared to its showy cousins.

Now, though, the shy little Calceocrinus balticensis gets a moment of exposure and formal admission to the roll call of life’s species.


Ausich, W.I. 1998. Phylogeny of Arenig to Caradoc crinoids (Phylum Echinodermata) and suprageneric classification of the Crinoidea. The University of Kansas Paleontological Contributions, n.s. 9, 36 pp.

Ausich, W.I. and Copper, P. 2010. The Crinoidea of Anticosti Island, Québec  (Late Ordovician to Early Silurian). Palaeontographica Canadiana 29, 157 pp.

Ausich, W.I., Wilson, M.A. and Vinn, O. 2012. Crinoids from the Silurian of western Estonia. Acta Palaeontologica Polonica 57: 613-631.

Wooster’s Fossils of the Week: A stromatoporoid-stromatolite combination (Upper Silurian of Saaremaa Island, Silurian)

July 22nd, 2012

There are two common fossil types that begin with “strom” and look roughly alike to the untrained eye. One is the stromatoporoid, which is a calcareous sponge, and the other is the stromatolite, which is a layered structure produced by photosynthetic bacteria. I hadn’t seen them together until our expedition to the Silurian of Estonia last summer. Wooster senior Nick Fedorchuk (’12) collected the specimen above at his outcrop of limestones and dolomites just above the Wenlock/Ludlow Boundary along Soeginina Cliff, Saaremaa. (In the rock sequence Richa Ekka is now studying.) We thought it was simply a stromatolite until he cut it to show that the base was a stromatoporoid.
“Stroma” is Greek for a bed or layer. Both stromatolites and stromatoporoids have horizontally laminated structures. The “lite” in stromatolite means rock, so a stromatolite is literally a “layered rock”. They are accretionary structures made by mostly cyanobacteria that collect and bind fine sediment into thin layers, usually in very shallow waters. Often the bacteria make their own calcareous cement for these laminae as a byproduct of photosynthesis. They’ve been doing this for a long time: the earliest known fossils are 3.5 billion-year-old stromatolites.

Stromatoporoids are very different. The “poroid” refers to their semi-porous skeletal layers, which are separated from each other by minuscule pillars. Their peak of abundance was in the Silurian and Devonian Periods, but they survived all the way up into the Cretaceous. They made significant reefs in the Paleozoic, often more common than the corals back then. We believe that they were a type of sponge (Phylum Porifera) with a thin layer of soft tissue on the exterior layer filter-feeding in the typical sponge manner.

Stromatolites are more common in sediments formed in very shallow, warm marine waters with elevated salinity; stromatoporoids liked more normal marine conditions. Finding the stromatolite on top of the stromatoporoid here means that either the environment changed between the two (shallowing, likely), or that the stromatoporoid was dislodged from more offshore waters during a storm and washed into a shallow lagoon, becoming a substrate for stromatolitic growth.

Curiously, there was a suggestion in 1990 by Kaźmierczak and Kempe that stromatoporoids ARE stromatolites. They pointed out that precipitation features in modern stromatolites can be very complex, producing features that resemble those of ancient stromatoporoids. This idea gained no traction, though, and most paleontologists are satisfied that these two types of “strom” have very different origins.


Akihiro, K. 1989. Deposition and palaeoecology of an Upper Silurian stromatoporoid reef on southernmost Gotland, Sweden. Geological Journal 24: 295-315.

Kaźmierczak, J. and Kempe, S. 1990. Modern cyanobacterial analogs of Paleozoic stromatoporoids. Science 250, no. 4985, pp. 1244-1248.

Lebold, J.G. 2000. Quantitative analysis of epizoans on Silurian stromatoporoids within the Brassfield Formation. Journal of Paleontology 74: 394-403.

Segars, M.T. and Liddell, W.D. 1988. Microhabitat analyses of Silurian stromatoporoids as substrata for epibionts. Palaios 3: 391-403.

Soja, C.M., White, B., Antoshkina, A., Joyce, S., Mayhew, L., Flynn, B. and Gleason, A. 2000. Development and decline of a Silurian stromatolite reef complex, Glacier Bay National Park, Alaska. Palaios 15: 273-292.

Vinn, O. and Wilson, M.A. 2010. Endosymbiotic Cornulites in the Sheinwoodian (Early Silurian) stromatoporoids of Saaremaa, Estonia. Neues Jahrbuch für Geologie und Paläontologie, Abh., v. 257: p. 13–22.

Analysis of a Rhuddanian (Llandovery, Lower Silurian) sclerobiont community in the Hilliste Formation on Hiiumaa Island, Estonia: a hard substrate-dwelling recovery fauna — An abstract submitted to the Geological Society of America for the 2012 annual meeting

July 19th, 2012

Editor’s note: The Wooster Geologists in Estonia this summer wrote abstracts for posters at the Geological Society of America Annual Meeting in Charlotte, North Carolina, this November. The following is from student guest blogger Jonah Novek in the format required for GSA abstracts:

Analysis of a Rhuddanian (Llandovery, Lower Silurian) sclerobiont community in the Hilliste Formation on Hiiumaa Island, Estonia: a hard substrate-dwelling recovery fauna

NOVEK, Jonah M., WILSON, Mark A., EKKA, Richa N., Department of Geology, The College of Wooster, Wooster, OH 44691 USA; AUSICH, William I., School of Earth Sciences, The Ohio State University, Columbus, OH 43210 USA; VINN, Olev, Department of Geology, University of Tartu, Ravila 14A, 50411 Tartu, Estonia

The Hilliste Formation on the island of Hiiumaa, western Estonia, is a Rhuddanian (Llandovery, Lower Silurian) sequence of limestones and shales.  It represents some of the earliest Silurian rocks on the paleocontinent of Baltica. The depositional system was tropical and shallow marine with tempestites indicated by overturned and broken corals and stromatoporoids. This unit contains a recovery fauna from the Ordovician Mass Extinction. Major taxa in the Hilliste Formation include crinoids, trilobites, bryozoans, corals, stromatoporoids, gastropods, and brachiopods. Sclerobiont communities (organisms that lived on or within hard substrates) have not yet been described from Rhuddanian faunas. The Hilliste Formation contains many encrusters and a few borings on skeletal substrates (primarily corals and crinoid stems). These sclerobionts include at least three kinds of crinoid holdfasts, cornulitids, sheet-like bryozoans, runner-type bryozoans, erect bryozoan holdfasts, and auloporid corals. Most if not all of these sclerobionts inhabited dead substrates. We studied the Hilliste Formation in a small quarry near the village of Hilliste on Hiiumaa. Numerous encrusted and bored specimens were collected for analysis of sclerobiont occurrences in this rare example of a Rhuddanian hard substrate community. These encrusters and borings, along with the macrofauna, have a distinct Late Ordovician aspect.

Stratigraphy and paleoenvironment of the Soeginina Beds (Paadla Formation, Lower Ludlow, Upper Silurian) on Saaremaa Island, Estonia — An abstract submitted to the Geological Society of America for the 2012 annual meeting

July 19th, 2012

Editor’s note: The Wooster Geologists in Estonia this summer wrote abstracts for posters at the Geological Society of America Annual Meeting in Charlotte, North Carolina, this November. The following is from student guest blogger Richa Ekka in the format required for GSA abstracts:

Stratigraphy and paleoenvironment of the Soeginina Beds (Paadla Formation, Lower Ludlow, Upper Silurian) on Saaremaa Island, Estonia

EKKA, Richa N., WILSON, Mark A., NOVEK, Jonah M., Department of Geology, The College of Wooster, Wooster, OH 44691 USA; VINN, Olev, Department of Geology, University of Tartu, Ravila 14A, 50411 Tartu, Estonia

The Soeginina Beds in the Paadla Formation on the island of Saaremaa, western Estonia, are a Lower Ludlow (Upper Silurian) sequence of dolostones, marls, and stromatolites. They represent rocks just above the Wenlock/Ludlow boundary, which is distinguished by a major disconformity that can be correlated to a regional regression on the paleocontinent of Baltica. We interpret the depositional environment of the Soeginina Beds as having been a hypersaline lagoon. Our evidence includes halite crystal molds, oscillation ripples, eurypterid fragments, stromatolites, ostracods, gastropods, Chondrites trace fossils, intraclasts and oncoids. Nautiloid conchs are common, probably because storm currents washed them in. We measured two sections of the Soeginina Beds at Kübassaare, eastern Saaremaa, western Estonia. The beds in one section are virtually horizontal; in the second they are steeply dipping, probably because of Pleistocene glacial ice overpressure. The beds begin with fine-grained dolostone and end with large, well-preserved domical stromatolites. The equivalent section at Soeginina Pank in western Saaremaa (about 86 kilometers away) has larger oncoids, branching coral fragments, and smaller stromatolites. It is also more heavily dolomitized. We interpret these differences as showing the western Soeginina Beds were deposited in slightly deeper, less saline waters than those in the east at Kübassaare.

Our last evening in Estonia

July 17th, 2012

TALLINN, ESTONIA–We had our final group dinner in the Olde Hansa restaurant. (Freezing at an outside table, I might add, but the food and company was excellent.) We leave for the USA at 5:00 a.m. tomorrow morning. Pictured above, just after our hearty dinner, are, from the left, Mark Peter, Jeff Thompson, Alyssa Bancroft, Mark Wilson, Jonah Novek, Bill Ausich, and Richa Ekka. It is a delightful team. We are already looking forward to our Ohio reunion.

Bill’s image of our last dinner in Tallinn. So much fun, especially after such a successful field trip. Note that it was a bit chilly!

Some wildflowers from the western islands of Estonia

July 17th, 2012

TALLINN, ESTONIA–This is a tradition on my field trips: a sampling of flowers we’ve seen. We are so fortunate to be working in such a beautiful place. It would be a pity not to share. I don’t have any identifications, so feel free to add them in the comments. [The above flower is Campanula persicifolia.]

(Actually I do know that this is a thistle and a Six-Spotted Burnet moth.)

This wild rose has raindrops from the first downpour we experienced at Undva Pank.

This geranium-like flower is one of the most common on Saaremaa.

This is the first time I’ve seen these little red stars.

These long yellow shoots were found only on the alvars on the Sarve Peninsula on Hiiumaa.

These look a bit like Mountain Laurel at home.

I have no idea what these lily-like flowers are. They are striking in the field.

A museum visit: Institute of Geology at Tallinn University of Technology

July 17th, 2012

TALLINN, ESTONIA–This morning Bill Ausich and I examined fossils in the collections of the Institute of Geology at Tallinn University of Technology (the cool exterior of which is shown above). The Chief Curator, Ursula Toom, generously came in from her vacation to show us some important Ordovician and Silurian crinoid specimens, as well as assemblages from the Lower Silurian throughout Estonia. We had an excellent time looking at gorgeous fossils in a classic museum. (We were here in 2009 with Rob McConnell and Palmer Shonk as well.)

It must be an interesting place if they bolt a rock up off the ground in front!

Bill is here looking at crinoids Ursula set aside for us to examine. Note his use of an iPad for taking notes and images, just like Wooster geologists did last year at Ohio State. Bill carries his entire pdf library with him on his iPad, and makes many annotated images of museum specimens.

Typical hall of cabinets in the Institute of Geology. Each set of drawers is on a mechanical device for closing the aisles to increase storage space.

This is a row of cabinets with one drawer opened. Note the use of a drawer partially opened underneath for support. (A rough experience once before I learned this trick …)

A typical drawer of specimens. These are newly collected from the Reinu Quarry by our friend and colleague Olev Vinn.

A specimen label. Unfortunately some are nearly indecipherable. Sometimes it is because a Russian worker was transliterating information into Latin letters. There is often an interesting mix between the Russian, Estonian and English languages. Fortunately Ursula and others can quickly translate for us!

We enjoyed working in the Institute of Geology collections very much. They are not only superbly organized, much of their content is listed (and even imaged) online. We saw many critical specimens, and Bill was able to borrow some important crinoids. Thank you to Ursula for her kindness and excellent assistance!


Goodbye Hiiumaa, Hello Tallinn!

July 16th, 2012

TALLINN, ESTONIA–Early this morning the Wooster/OSU geologists left the Estonian island of Hiiumaa on the ferry from the port of Heltermaa (58° 52′ N, 23° 3′ E; seen above). After a 90-minute ferry ride, we landed on the Estonian mainland. We looked briefly at some quarries near Pusku south of Haapsalu (the limestones were too dolomitized for our liking) and then made the two-hour drive to Tallinn. After negotiating the most miserable intersection in Europe (which is under perpetual construction), we at last gassed up our cars and turned them in to the Sixt rental agency. There is nothing quite like the feeling of returning vehicles safely after such a long trip — it is only topped by returning students safely to their homes, of course! We are now in Tallinn for two nights. Bill and I have an appointment with the chief curator at the Institute of Geology in Tallinn as our students take a day off in the city. We are all staying at the St. Barbara Hotel near Old Town — a place beloved by many past Wooster students (depending on whether we have drunk Russians serenading us from the street all night.) I like that the building stone is a beautiful local Ordovician limestone.

The coiled-and-ribbed fossil mystery deepens on Hiiumaa

July 15th, 2012

KÄINA, ESTONIA–It has been a rainy day on the Estonian island of Hiiumaa. The Wooster geologists stayed inside most of the day to work on their Geological Society of America abstracts. Bill Ausich and his Ohio State University team, though, returned to the Hilliste Quarry and continued to collect fossils. To our great surprise, they picked up two more specimens of that strange planispirally-coiled shell that Richa found on July 13. One is shown above, and the “venter” view is shown below. All specimens were found in a yellowish unit that matches the matrix in the fossils (although we’ll check in the lab to make certain). These new finds reduce the chances that the fossils are a product of “site contamination” in which a visitor discards specimens from a previous trip, often to make room for new ones. That is still a possibility, but an increasingly remote one.

So what are these? They look very much like Mesozoic ammonites, all the way down to deflections of ribs along the periphery as you might be able to see above. (Specimen photography in a hotel room has its challenges.) The earliest ammonoids, the larger group that contains ammonites, appear in the Devonian (the period after the Silurian), so it is unlikely we are looking at that group. They are certainly mollusks, though, so most likely gastropods (snails) or nautiloids. Coincidentally enough, Wooster alumnus James St. John has a webpage with a photograph of a coiled, ribbed nautiloid known as Graftonoceras, which you will note has many similarities with our mystery critter. The specimen he photographed is in the museum at, of all places, Ohio State!

The last holdouts of pagan Europe

July 14th, 2012

KÄINA, ESTONIA–The little island of Muhu between Saaremaa and the Estonian mainland, had a large prehistoric population — much larger than it has today. The Muhu Estonians built a large fort of stone heaps near the western coast opposite Saaremaa so that they could control the traffic and trade through the Small Strait. The remains of that fortification are seen above. In January 1227, Teutonic Crusaders cornered the last of the pagan Estonians in this stronghold. (They were, in fact, among the last pagans in all of Europe.) Reports say that 20,000 soldiers besieged 2500 Estonian warriors for six days here. All the Estonians were killed save one, who escaped by pretending to be a victorious crusader. Most of the stones of the fort were removed to build the causeway between Muhu and Saaremaa, but the site remains as a ring of earthen walls and a stone monument (below) marking the bloody battlefield.

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