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Wooster’s Fossils of the Week: An Upper Ordovician cave-dwelling bryozoan fauna and its exposed equivalents

July 3rd, 2015

1 Downwards 063015This week’s fossils were the subject of a presentation at the 2015 Larwood Symposium of the International Bryozoology Association in Thurso, Scotland, last month. Caroline Buttler, Head of Palaeontology at the National Museum Wales, Cardiff, brilliantly gave our talk describing cryptic-and-exposed trepostome bryozoans and their friends in an Upper Ordovician assemblage I found years ago in northern Kentucky. They were the subject of an earlier Fossil of the Week post, but Caroline did so much fine work with new thin sections and ideas that they deserve another shot at glory. We are now working on a paper about these bryozoans and their borings. Below you will find the abstract of the talk and a few key slides to tell the story.


Trepostome bryozoans have been found as part of an ancient cave fauna in rocks of the Upper Ordovician (Caradoc) Corryville Formation exposed near Washington, Mason County, Kentucky.

Bryozoans are recognized as growing from the ceiling of the cave and also from an exposed hardground surface above the cave. Multiple colonies are found overgrowing one another and the majority are identified as Stigmatella personata. Differences between those growing upwards and those growing down from the roof have been detected in the limited samples.

The colonies have been extensively bored, these borings are straight and cylindrical. They are identified as Trypanites and two types are recognised. A smaller variety is confined within one colony overgrowth and infilled with micrite. In thin section it is observed that the borings follow the lines of autozooecial walls and do not cut across. This creates a polygonal sided boring, suggesting that the colonies were not filled with calcite at the time of the boring. The second variety has a larger tube size and its infilling sediment has numerous dolomite rhombs and some larger fossil fragments including cryptostomes, shell and echinoderm pieces. These cut through several layers of overgrowing bryozoans. Some of the borings contain cylindrical tubes of calcite similar to the ‘ghosts’ of organic material described by Wyse Jackson & Key (2007).

Very localised changes in direction of colony growth due to an environmental effect are seen.

Bioclaustration in these samples provides evidence for fouling of the colony surface, indicating that the bryozoans overgrew unknown soft-bodied organisms.


Wyse Jackson, P. N., and M. M. Key, Jr. (2007). Borings in trepostome bryozoans from the Ordovician of Estonia: two ichnogenera produced by a single maker, a case of host morphology control. Lethaia. 40: 237-252.

2 Title 0630153 Location 0630154 Strat position 0630155 hdgd up 0630156 hdgd down 0630157 Growth up 0630158 Growth down 0630159 Stigmatella 06301510 Cartoon 06301511 Boring A 06301512 Boring B 06301513 Ghosts explanation14 Ghosts 06301515 Overgrowths 06301516 Further questions 063015

Link to posts from Wooster Geologists in the United Kingdom in June 2015

June 29th, 2015

11 Mae Meredith Filey BriggI spent 25 days in England, Scotland and Wales this month, 12 of them with these two happy Senior Independent Study students, Mae Kemsley (’16) and Meredith Mann (’16) — dubbed “Team Yorkshire”. We had to delay our blog posts until today. You can see all of them by clicking the UK2015 tag. It was a spectacular expedition. Thanks again to Paul Taylor, Jen Loxton, Joanne Porter, and Tim Palmer for the parts they played in this adventure. Thank you as well to Mae and Meredith who were not only sharp field paleontologists, they were great companions as well. They are shown above on the tip of Filey Brigg in North Yorkshire. (N54.21560°, W00.25842°; Google Earth image below. Cool study site!)

Screen Shot 2015-06-29 at 11.47.54 AM

Shikotan Island Tree Ring Chronology

June 26th, 2015

Guest blogger: Xiangyu Li

As one of the most militarized islands, because of the dispute between Japan and Russia (the Kuril Islands dispute), Shikotan Island has remained a mystery to the world of tree rings and climate studies until now.

1447443_original Figure 1. Deserted Russian tank on Shikotan island (from

Shikotan Island directly on the margin between the Pacific and Eurasian Plates. This special location indicates that this island is greatly influenced by geological factors, such as tsunami, earthquake, and large changes in climate (see photos below).


Figure 2. Big crack after the Shikotan Island earthquake of 1994.

see ice

Figure 3. Shikotan Island surrounded by sea ice (Wikipedia).

In order to better define the climate history of the Island in 2014 Russian geologist E. Dolgova and M. Alexandrin collected core samples from larch, spruce and fir trees on Shikotan islands.


Figure 4. Location of  collection sites of all Shikotan cores. The island is approximately 30 km long (image from GooglEarth).

The purpose of this research is to find the climate information from these cores. These cores are properly sanded and marked. Forty six cores were collected from Shikotan Island and 20 were used to construct the chronology for this island.


Figure 5. Family picture of all cores in the chronology

As we can see, some cores are less than 100 years old and some cores can be dated back to 18th century. Despite the age difference these 20 cores correlate with each other well.The purpose of this research is to extract climate information from these cores. These cores are properly sanded and marked by year. Forty six cores were collected from Shikotan Island and 20 were used to construct the chronology for this island.

standardized tree ring indices

The tree ring width can be influenced by various factors other than climate signal, such as the growth trends and natural competition. The growth trends is the major factor that we want to eliminate. The above picture shows us the standardized tree ring measurements by removing the growth trends. The blue line is the sample size. This is the first tree-ring record from Shikotan Island.

Final copy

The standardized ring width chronology has a slight downward trend since 1900. I compared the standardized tree ring data with the meteorological data from Nemuro station (130 years long). It appears that September temperature has a strong negative correlation with the tree ring width. The correlation is 0.43, which is far above the 99% confidence level. The reason for such negative correlation is unknown, however, it may be related to changes in sea ice extent.

In addition to examining the correlation between ring width and temperature, I focused on the possible relationship between tree ring width and natural hazards, such as tsunami and earthquake.

After looking at the tsunami occurrence data from NOAA, I found that sometimes tsunami corresponded with a year of rapid growth. For example, in 1963, there are two tsunamis on Shikotan Island and some cores has a bigger growth this year compared to the year before. Photos below are two cores that show the correlation with the tsunami record and there are more cores have a correlation with the tsunami record. This relationship is under investigation.


Figure 6. Core K02E5B


Figure 7 K02E7B

Last day of fieldwork in England: A working quarry and another great unconformity

June 26th, 2015

1 Doulting quarry sawBRISTOL, ENGLAND (June 26, 2015) — Tim Palmer has a professional interest in building stones, and a passion for sorting out their characteristics and historical uses. He thus has many contacts in the stone industry, from architects to quarry managers. This morning we visited the Doulting Stone Quarry on the outskirts of Doulting near Shepton Mallet in Somerset. Here a distinctive facies of the Jurassic Inferior Oolite is excavated for a variety of purposes. The rock has a lovely color, is relatively easy to work, and is durable. Above is a quarry saw that cuts out huge blocks from the natural exposure.

2 Thalassinoides layer DoultingSuch sawing produces great cross-sections for geologists to examine. We were particularly interested in that light-colored unit above with the irregular top and dark sediment-filled holes. The holes are part of a network of Thalassinoides burrows (tunnels made by Jurassic crustaceans) and reduce the value of the rock as a building stone. There is thus lots of it laying around the quarry yard for study.

3 Pinnid likely Trichites cross section DoultingOne impressive fossil exposed by the sawing is this pinnid bivalve, probably Trichites.

4 Burrow fill sediments DoultingThe Thalassinoides burrows are filled with a poorly-cemented sediment. It is full of little fossils, so we collected a bag of it for microscopic examination. It may give us clues as to what communities lived on the surface of this burrowed unit when it was part of the Jurassic seafloor.

5 shaping saw DoultingWe had a tour of the quarry shops, which included seeing these giant rock saws in action. Many of the saws are controlled by computers, so elaborate cuts can be made.

6 Medieval stone breaking marksThis rock has been quarried since Roman times, so there is over 2000 years of stone working here. The quarry owner set aside this rock face which shows chisel marks made in Medieval times. Wooden wedges were jammed into chiseled channels and then pulled over days to eventually crack the stone free.

7 Tedbury Camp wavecut surface along strikeAfter the quarry visit, Tim Palmer and I tromped through the woods and eventually found (with the help of several locals) an exposure known as Tedbury Camp. It is another Jurassic-on-Carboniferous unconformity like we saw at Ogmore-By-Sea earlier in the week. A century ago quarry workers cleared off this surface of Carboniferous limestone. It is a wave-cut platform on which Jurassic sediments (the Inferior Oolite) were deposited. The surface has many geological delights, including faults, drag folds, differentially-weathered cherts and carbonates, and Jurassic borings and encrusters. Beautiful.

8 wavecut surface foldingIn this view of the surface you may be able to see the odd folding of the dark chert layers in the right middle of the image. These seem to be drag folds along a fault. They clearly predate the Jurassic erosion of the limestone surface. The overlying Jurassic can be seen in the small outcrop on the left near Tim.

9 section view of wavecut surfaceIn this cross-section of the erosional surface you can clearly see we’re working with an angular unconformity.

10 filled borings wavecutTrypanites borings are abundant across this surface, most filled with lighter Jurassic sediment. There are other borings here too that deviate from the straight, cylindrical nature of Trypanites.

11 curved borings wavecutI don’t know yet how to classify these curved borings. They resemble Palaeosabella.

12 Encrusting bivalve wavecutHere is a Jurassic bivalve attached to the Carboniferous limestone at the unconformity. Most of the encrusters have been eroded away.

13 Tim on wavecut platformThere are many possibilities for further study of the Tedbury Camp unconformity. This was a productive site for our last field visit in England this year. Thank you very much to Tim Palmer, seated above, for his expertise, great companionship, and generosity with his time. It was a reminder of how much fun we had together in the field twenty years ago.

My month of geology in the United Kingdom has now come to an end. My next two days will be devoted to packing up and making the long train and then plane flights home. What a wonderful time I had, as did my students on the earlier part of the trip, Mae Kemsley and Meredith Mann. Thank you again to Paul Taylor for his work with us in Scarborough. I am very fortunate with my fine British friends.

For the record, the important locality coordinates from this trip —

GPS 089: Millepore Bed blocks N54.33877°, W00.42339°

GPS 090: Spindle Thorn Member, Hundale Point N54.16167°, W00.23326°

GPS 091: Robin Hood’s Bay N54.41782°, W00.52501°

GPS 092: Northern limit of Speeton Clay N54.16654°, W00.24567°

GPS 093: Northern limit of Red Chalk N54.15887°, W00.22261°

GPS 094: South section Filey Brigg N54.21674°, W00.26922°

GPS 095: North section Filey Brigg N54.21823°, W00.26904°

GPS 096: Filey Brigg N54.21560°, W00.25842°

GPS 097: D6 of Speeton Clay N54.16635°, W00.24520°

GPS 098: C Beds of Speeton Clay N54.16518°, W00.24226°

GPS 099: Lower B Beds of Speeton Clay N54.16167°, W0023326°

GPS 100: Possible A Beds of Speeton Clay N54.16129°, W00.23207°

GPS 101: A/B Beds of Speeton Clay N54.16035°, W00.22910°

GPS 102: C7E layer of Speeton Clay N54.16447°, W00.24043°

GPS 103: Lavernock Point N51.40589°, W03.16947°

GPS 104: Triassic deposits, Ogmore-By-Sea N51.46543°, W03.64094°

GPS 105: Sutton Stone Unconformity N51.45480°, W03.62609°

GPS 106: Sample of lowermost Sutton Stone N51.45455°, W03.62545°

GPS 107: Nash Point N51.40311°, W03.56212°

GPS 108; Devil’s Chimney N51.86402°, W02.07905°

GPS 109: Fiddler’s Elbow N51.82584°, W02.16541°

GPS 110: Doulting Stone Quarry N51.18993°, W02.50245°

GPS 111: Tedbury Camp unconformity N51.23912°, W02.36515°


Wooster’s Fossils of the Week: An encrusted bivalve external mold from the Upper Ordovician of Indiana

June 26th, 2015

1 Anomalodonta gigantea Waynesville Franklin Co IN 585I love this kind of fossil, which explains why you’ve seen so many examples on this blog. We are looking at an encrusted external mold of the bivalve Anomalodonta gigantea found in the Waynesville Formation exposed in Franklin County, Indiana. I collected it many years ago as part of an ongoing study of this kind of preservation and encrustation.
2 Anomalodonta gigantea Waynesville Franklin Co IN 585 annotatedTo tell this story, I’ve lettered the primary interest areas on image above. First, an external mold is an impression of the exterior of an organism. In this case we have a triangular clam with radiating ribs in its shell. The exterior of the shell with its ribs was buried in sediment and the shell dissolved, leaving the basic impression above. It is a negative relief. Please now refer to the letters for the close-up images below.

3 Bryo Anomalodonta gigantea Waynesville Franklin Co INA. At the distal end of the bivalve mold is what looks at first to be the original shell. It is calcitic, though, and we know this bivalve had an aragonitic shell. A closer look shows that this is actually the attaching surface of an encrusting bryozoan that bioimmured the original bivalve shell, which has since dissolved away. This smooth surface is the bryozoan underside; we see the characteristic zooecia (tubes holding the individual zooids) only when this surface is weathered away.

4 Borings Anomalodonta gigantea Waynesville Franklin Co INB. These tubular objects are infillings of borings (maybe Trypanites)that were cut into the original aragonitic shell of the bivalve. The tunnels of the borings were filled with fine sediment, and then the shell dissolved away, leaving these casts of the borings.

5 Inarticulate scar Anomalodonta gigantea Waynesville Franklin Co INC and D. In the middle of the external mold is this curious circular feature (C) mostly surrounded by a bryozoan (D). There was at one time a circular encruster, likely an inarticulate brachiopod like Petrocrania, that sat directly on the external mold surface. The bryozoan colony grew around but not over it because it was alive and still opening and closing its valves for feeding. The bryozoan built a vertical sheet of skeleton around it as a kind of sanitary wall. You may be able to see the other three or four structures in the top image showing brachiopod encrusters that left the building. This is an example of fossils showing us a living relationship, even if one is not longer preserved.

This fossil and its sclerobionts (hard substrate dwellers) show us that soon after the bivalve died its aragonitic shell dissolved away, leaving as evidence the external mold in the sediment, the bioimmuring bryozoan, and the boring casts. Very soon thereafter bryozoans and brachiopods encrusted the available hard substrate. This is a typical example of early aragonite dissolution on the sea floor during a Calcite Sea interval.


Palmer, T.J. and Wilson, M.A. 2004. Calcite precipitation and dissolution of biogenic aragonite in shallow Ordovician calcite seas. Lethaia 37: 417-427.

Taylor, P.D. 1990. Preservation of soft-bodied and other organisms by bioimmuration—a review. Palaeontology 33: 1-17.

Taylor, P.D. and Wilson, M.A. 2003. Palaeoecology and evolution of marine hard substrate communities. Earth-Science Reviews 62: 1-103.

Wilson, M.A., Palmer, T.J. and Taylor, P.D. 1994. Earliest preservation of soft-bodied fossils by epibiont bioimmuration: Upper Ordovician of Kentucky. Lethaia 27: 269-270.

Wooster Geologist in England (again)

June 25th, 2015

1 old quarry face cotswoldsBRISTOL, ENGLAND (June 25, 2015) — Our little geological exploration of southern Britain now passes into England. Tim Palmer and I crossed the River Severn and drove to the Cotswolds to examine old quarry exposures and Medieval stonework. We are parked above in Salterley Quarry near Leckhampton Hill.

2 devils chimney leckhamptonOur theme again is Jurassic. At Leckhampton Hill we examined exposures of the Middle Jurassic Inferior Oolite. It is not, of course, inferior to anything in the modern sense. The name, originally from William Smith himself, refers to its position below the Great Oolite. This is Devil’s Chimney, a remnant of stone left from quarrying in the 19th Century.

3 fiddlers elbow hdgd and Pea GritWe stopped along a bend in a Cotswold road called Fiddler’s Elbow and found an old carbonate hardground friend in the Inferior Oolite. Borings are evident in this flat, eroded surface. Next to the hammer are pieces of the Pea Grit, a coarser facies. I want to examine the grains for microborings and encrusters.

4 Dogrose leckhamptonThis is the gorgeous dog-rose (Rosa canina, not surprisingly), which is common in the Cotswolds. It is the model for the Tudor Rose in heraldry.

5 Fiddlers elbow orchidsThese tall orchids were also abundant near our outcrops.

6 fiddlers elbow orchids closeA closer view of the orchids. When I learn the name for this plant, I’ll amend this post.

7 Painswick church and yewsAt the end of the day we stopped by St. Mary’s Church in Painswick, with its distinctive churchyard and variety of building stones. The sculptured trees are English yews.

8 Tim and Painswick gravestonesThe gravestones date back to the early 18th Century, with older ledger stones inside the church.

9 Painswick church pyramid markerThe unique pyramidal tomb of the stonemason John Bryan (1716-1787). He was apparently responsible for most of the 18th Century ornate monuments in this churchyard.

10 copper markers killing lichen PainswickMany of the gravestones have copper plates affixed to their upper faces. The rain washes copper ions out of the metal and over the limestone, killing the lichens and other encrusting organisms. This leaves the lighter patch of bare limestone. Somewhere in this is a study of microbiome ecological gradients!

11 St Marys church Painswick shot divot 1643The Painswick church was the site of a 1643 battle during the English Civil War. There are numerous bullet and shot marks on the exterior stones. Tim commented on the remarkable resilience of this stone to stay coherent after almost 400 years of weathering of these pits.

National Museum Wales and its new dinosaur

June 25th, 2015

1 National Museum WalesBRIDGEND, WALES (June 25, 2015) — On our last day in Wales, Tim had an errand at the National Museum Wales in Cardiff. We took the opportunity to visit their new dinosaur exhibit with the skeleton that had been collected from an outcrop we visited earlier in the week at Lavernock Point.

2 Dino displayThe exhibit is well done. The fossil skeleton represents the first carnivorous dinosaur found in Wales, and one of the earliest Jurassic dinosaurs found anywhere.

3 Lavernock dino bonesHere are some of the bones in lower Lias limestone.

4 Dino site museum imageThis is a museum photo of the dinosaur collecting site. You may recognize the place from a previous post in this blog.

Cardiff city hallThis is Cardiff City Hall. I loved the early 20th Century architecture in this Cardiff district, but we didn’t have time to explore. It is now off to southern England for more fieldwork.

A great unconformity in South Wales

June 24th, 2015

1 Dinantian Sutton unconformity wide viewBRIDGEND, WALES (June 24, 2015) — Today Tim Palmer and I visited a famous unconformable rock plane in South Wales. I last saw it thirty years ago, when I knew a lot less about eroded, bored and encrusted surfaces. It is an unconformity between a Carboniferous limestone (High Tor Limestone, Dinantian in age) and an overlying Jurassic limestone (Sutton Stone, Hettangian, Lower Jurassic) exposed on the coast near Ogmore-By-Sea. It was most thoroughly described in 2004 by Johnson and McKerrow (Palaeontology 38: 529-541). You can see it as the surface above, with the Jurassic rocks on top of it to the right. (I know, gray rocks on gray rocks. It takes close examination to tell them apart after they both have been subjected to coastal weathering.)

2 Dinantian Sutton close viewHere is a closer view with part of the Lower Jurassic Sutton Stone broken away to show fresh material. (We didn’t do this, despite the guilty-looking hammer. The hammer is Paul Taylor’s, by the way. Thanks, Paul!) Note the pebbles in the Sutton Stone. They are made of the Carboniferous limestone beneath. Classic unconformity.

3 Dinantian Sutton borings wide viewThe Carboniferous limestone is punctured by numerous small borings (Trypanites) drilled by filter-feeding worms of some kind when the Early Jurassic sea covered this surface. They are the clusters of small black dots shown above.

4 Dinantian Sutton borings closeIn this closer view of the borings you can see that they are filled with a lighter Jurassic sediment. The openings have been somewhat enlarged by weathering.

5 Dinantian Sutton reliefThis erosion surface shows some relief, probably formed by cobbles and pebbles washing over it during the Early Jurassic. This matches what we see on modern wave-cut rocky platforms.

6 Triassic Dinantian unconformityOn the same stretch of shoreline there is a small section where Triassic wadi deposits cut down into the Carboniferous limestone — another even more dramatic unconformity, but without marine fossils.

7 Triassic wadi deposits Ogmore by seaComing from a desert myself, I have an affinity for wadi sediments. They are coarse, angular and poorly sorted. These grains are entirely from the underlying Carboniferous limestone. They were likely generated from steep rocky canyons through which intermittent streams flowed.

8 Nash Point viewAt the end of the day Tim and I visited Nash Point, again on the coast of South Wales. Here the Lias is brilliantly (and dangerously!) exposed as a series of alternating limestones and shales.

9 Nash Point CliffWe didn’t get too close to these unstable cliffs. The limestone blocks fall often as the interbedded soft shales holding them in place weather away.

10 Nash PointA view of Nash Point at low tide. Tim always wears that red jacket, so he’s easy to spot. (Classic Redcoat!)

11 Nash Point cobblesWe didn’t find much to paleontologically interest us at this last outcrop, but it was beautiful on another stunning coastal day. These cobbles, all made of Lias limestones, are pretty to look at, but tiresome to walk through. We were ready for a slow dinner after this excellent day.


Wooster Geologist in Wales

June 23rd, 2015

1 Triassic Lavernock Point Penarth GroupBRIDGEND, WALES (June 23, 2015) — My train journey yesterday was successful. It was close, but I made the four tight connections and arrived in Aberystwyth, Wales, from Thurso, Scotland, on schedule. It took 15 hours. My friend Tim Palmer was there to greet me as I stumbled out of my carriage. I went from rainy, cold Scotland to warm and sunny Wales. The top image is of the Triassic/Jurassic transition at Lavernock Point in south Wales (see below).

2 Tim Caroline houseMy first Welsh night was with Tim in his great country home (with is wife Caroline) on the outskirts of Aberystwyth. It is called The Old Laundry because of its function on a previous manorial estate. I had my best sleep here for the entire trip. Quiet and beautiful.

4 Talley Abbey ruinsOne of Tim’s passions is the study of building stones in England and Wales. As we drove to southern Wales for our geological work, we stopped by interesting stone structures, including the ruins of Talley Abbey, a 12th Century monastery.

5 Tim at Talley AbbeyTim is here examining the dressing stones on a corner of this pillar in the Talley Abbey ruins. I learned that these dressings are usually made of stone that can be easily shaped, is attractive, and will hold sharp edges. In many cases these are called “freestones”.

6 Lower Lias Lavernock dinosaur siteOur first geological stop was at Lavernock Point on the southern coast of Wales. We looked here at the boundary section between the Upper Triassic and Lower Jurassic (Lias). In this view we see an alternating sequence of limestones (buff-colored) and shales (dark gray) of the lower Lias. These are marginal marine units with oysters and ammonites. On the left side of the image you can see a broad niche cut back into the cliff. This is the site where the first carnivorous dinosaur in Wales was recently excavated. It is also one of the oldest Jurassic dinosaurs since it was discovered just above the Triassic/Jurassic boundary. More on this dinosaur later.

7 Lower Lias pseudo mudcracksWe wandered across broad intertidal wave-cut platforms at Lavernock Point looking at the limestones and shales of the lower Lias. I was intrigued by these features on some bedding planes. They are not desiccation cracks, but rather some combination of jointing and weathering.

8 Liostrea hisingeri LavernockThe oyster Liostrea hisingeri is very common in this part of the Lias. In the limestones it is sectioned by erosion, resulting in these shelly outlines.

9 Liostrea hisingeri shale LavernockWhen Liostrea hisingeri is present in the shales, it is preserved three dimensionally.

10 Tonypandy street viewAfter our geologizing was done for the day, Tim and I drove up into the Rhondda Valleys just north of our hotel. This was at one time a very busy coal mining and industrial region, but the mines are closed and most of the heavy industry has moved elsewhere. Above is a view down a street in Tonypandy, one of the more famous towns of The Valleys. There were massive riots here in 1910 which eventually a minimum wage for miners in 1912.

Another long train journey through Great Britain

June 22nd, 2015

1 Thurso stationTHURSO, SCOTLAND (June 22, 2015) — Back to the trains today as I leave for a long journey south through Scotland and central England and then west to Aberystwyth, Wales, to spend some quality field time with my friend Tim Palmer. Our goal will be to explore sites for potential Independent Study projects.

4 Screen Shot 2015-06-24 at 8.51.34 PMLots of time on the trains coming up on this trek. Even more if I miss a connection. British train journeys often include only a few minutes to change trains at a station. This is not always a straightforward task.

2 Thurso track endsThe tracks at Thurso are the most northerly extension of the British rail system.

3 Tracks from ThursoSouth I go. There is no other direction!

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