Archive for February 23rd, 2014

Wooster’s Fossil of the Week: An interlocking rugose and tabulate coral (Devonian of Michigan)

February 23rd, 2014

Hexagonaria percarinata colony viewThis beautifully polished fossil looks like half of an antique bowling ball. Normally I hate polished fossils because the external details have been erased, but in this case the smooth surface reveals details about the organisms and their relationship. We have here a large colonial rugose coral with a smaller tabulate coral embedded within it. The specimen is from the Devonian of Michigan. It may look familiar because it is a large “Petoskey Stone“, the state stone (not fossil!) of Michigan. The large rugose coral is Hexagonaria percarinata (Sloss, 1939).
Hexagonaria percarinata close view 585In this closer view you can see the multiple star-like corallites of this coral. Each corallite held a tentacular feeding polyp in life. The radiating lines are thin vertical sheets of skeleton called septa. The corallites in this type of coral shared common walls and nestled up against each other as close as possible. In the lower center of the image you can see a very small corallite that represents a newly-budded polyp inserting itself as the colony grew. If rugose corals were like modern corals (and they probably were), the polyps were little sessile benthic carnivores catching small passing organisms with a set of tentacles. They may also have had photosymbionts to provide oxygen and carbohydrates through photosynthesis.
Tabulate coral intergrown with HexagonariaIn the midst of the rugose coral is this irregular patch with another type of coral: a tabulate coral distinguished by numerous horizontal partitions in its corallites (and no septa). It was likely a favositid coral, sometimes called a “honeycomb coral”. It was clearly living in the rugosan skeleton and not pushed into it by later burial. Note, though, the ragged boundary between the two corals. The rugose coral has the worst of it with some corallites deeply eroded. What seems to have happened is that the rugose coral had an irregular opening in its corallum (colonial skeleton) after death and the tabulate grew within the space, eventually filling it. The tabulate likely stuck out far above the rugose perimeter, but the polishing shaved them down to the same level. This is thus not a symbiotic relationship but one that happened after the death of the rugose coral.
Stumm, Erwin C   copyThe rugose coral species, Hexagonaria percarinata, was named in 1939 by Laurence Sloss, a famous sedimentary geologist with an early start in paleontology, but it is best known through the research of Erwin Charles Stumm (1908-1969; pictured above). Stumm was at the end of his life a Professor of Geology and Mineralogy and the Curator of Paleozoic Invertebrates in the Museum of Paleontology at the University of Michigan. Stumm grew up in California and then moved east for his college (George Washington University, ’32) and graduate (PhD from Princeton in 1936) education. He taught geology at Oberlin College up the road for ten years, and then moved to Michigan to start as an Associate Curator and Assistant Professor. I knew his name because in 1967 he was President of the Paleontological Society. He is said to have been a dedicated teacher of undergraduates and effective graduate advisor. It is fitting that his name is connected to such a popular fossil as Hexagonaria percarinata.

References:

Sloss, L. 1939. Devonian rugose corals from the Traverse Beds of Michigan. Journal of Paleontology 13: 52-73.

Stumm, E.C. 1967. Growth stages in the Middle Devonian rugose coral species Hexagonaria anna (Whitfield) from the Traverse Group of Michigan. Contributions from the Museum of Paleontology, The University of Michigan 21(5): 105-108.

Stumm, E.C. 1970. Corals of the Traverse Group of Michigan Part 13, Hexagonaria. Contributions from the Museum of Paleontology, The University of Michigan 23(5): 81-91.