Wooster’s Fossil of the Week: A receptaculitid (Middle Ordovician of Missouri)

September 18th, 2011

This week’s fossil is a long-standing paleontological mystery. Above is a receptaculitid from the Kimmswick Limestone (Middle Ordovician) near Ozora, Missouri. I think I found it on a field trip with Frank Koucky in the distant mists of my student days at Wooster, but so many outcrops, so many fossils …

Below is a nineteenth century illustration of a typical receptaculitid fossil. They are sometimes called “sunflower corals” because they look a bit like the swirl of seeds in the center of a sunflower. They were certainly not corals, though, or probably any other kind of animal. Receptaculitids appeared in the Ordovician and went extinct in the Permian, so they were confined to the Paleozoic Era. Receptaculitids were bag-like in form with the outside made of mineralized pillars (meroms) with square or diamond-shaped heads. The fossils are usually flattened disks because they were compressed by burial. You may notice now that the fossil at the top of this post is a mold of the original with the dissolved pillars represented by open holes. (Paleontologists can argue if this is an external or internal mold.)So what were the receptaculitids? When I was a student we called them a kind of sponge, something like a successor of the Cambrian archaeocyathids. In the 1980s a convincing case was made that they were instead a kind of alga of the Dasycladales. Now the most popular answer is that they belong to that fascinating group “Problematica”, meaning we have no idea what they were! (Nitecki et al., 1999). It’s those odd meroms that are the problem — they appear in no other known group, fossil or recent.

I find it deeply comforting that we still have plenty of fossils in the Problematica. We will always have mysteries to puzzle over.
Another Wooster receptaculitid specimen, this time seen from the underside showing side-views of the meroms.
Diagram of a receptaculitid in roughly life position showing its inflated nature and pillar-like meroms. From Dawson (1880, fig. 25): a, Aperture (probably imaginary here). b, Inner wall. c, Outer wall. n, Nucleus, or primary chamber. v, Internal cavity.

Finally, this is what a typical receptaculitid looks like in the field (Ordovician of Estonia). Note that nice sunflower spiral of the merom ends.

References:

Dawson, J.W. 1880. The chain of life in geological time: A sketch of the origin and succession of animals and plants. The Religious Tract Society, 272 pages.

Nitecki, M.H., Mutvei, H. and Nitecki, D.V. 1999. Receptaculitids: A Phylogenetic Debate on a Problematic Fossil Taxon. Kluwer Academic/Plenum, 241 pages.

6 Responses to “Wooster’s Fossil of the Week: A receptaculitid (Middle Ordovician of Missouri)”

  1. Louis Varricchioon 09 May 2012 at 8:52 am

    Hello: I am writing a newspaper article about fossil recepticulitids found in the Lake Champlain Valley of Vermont. I wonder if I may have permission to reprint your photo (above) with credit? Hope to hear from you very soon. Thank you!

  2. Mark Wilsonon 09 May 2012 at 9:02 am

    You certainly may! All my photos here and on my Wikimedia page

    http://commons.wikimedia.org/wiki/User:Wilson44691

    are public domain. You may use them as you wish. You may find higher resolution versions on Wikimedia and Wikipedia.

  3. Kiton 05 Jul 2013 at 10:42 am

    So if the thinking is that they are not animals, then could it be a plant…?

  4. Mark Wilsonon 05 Jul 2013 at 11:00 am

    Some kind of algae is most likely, Kit, but the debates continue.

  5. Jesse Garnett Whiteon 13 Aug 2013 at 1:27 pm

    Found some good specimens of these in ornamental building facing Tyndall Stone in Calgary, Canada.

    JGW

  6. Curtis Raddatzon 10 Dec 2013 at 11:32 am

    I have been finding really nice fossils of these in Northeast Iowa this last year.

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